Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet

Twelve species of the subgenus Vestiplex (Diptera, Tipulidae) were previously known to occur in Tibet (= Xizang), China. Here, six species are described and illustrated as new to science: Tipula (Vestiplex) bucera sp. nov., Tipula (Vestiplex) magatama sp. nov., Tipula (Vestiplex) motuoensis sp. nov., Tipula (Vestiplex) nayogabuensis sp. nov., Tipula (Vestiplex) platyphylla sp. nov. and Tipula (Vestiplex) uncinella sp. nov. The following three species are redescribed: Tipula (Vestiplex) himalayensis Brunetti, 1911, Tipula (Vestiplex) nigroapicalis Brunetti, 1911 and Tipula (Vestiplex) zayulensis Alexander, 1963. A key to the species of Tipula (Vestiplex) from Tibet is presented.


Introduction
The tipulid subgenus Tipula (Vestiplex) was erected by Bezzi (1924) for the species Tipula cisalpina Riedel, 1913 andT. hemapterandra Bezzi, 1924. The subgenus currently includes 178 known species and subspecies worldwide, which are distributed in the Holarctic (112 taxa) and the Oriental Realms (76 taxa) (Oosterbroek 2021). To date, 76 species are known from China, including 12 species from Tibet Autonomous Region (hereafter abbreviated as Tibet). Most species of this subgenus have variegated wings, while some species have reduced wings. The male tergite 9 commonly forms a shallowly concave sclerotized saucer or can be longitudinally divided into two parts, some species have two pubescent fi nger-like arms. The female ovipositor is commonly with saerrated outer margin (Alexander 1935(Alexander , 1965Savchenko 1964;Alexander & Byers 1981;Starkevich & Podenas 2011;Starkevich 2012;Men et al. 2017).
The currently described species of the subgenus Vestiplex are prevalent in the Holarctic and Oriental zoogeographical regions. The majority of the species are related to mountain systems (Alps, Pyrénées, Caucasus and Himalayas), at altitudes up to 4500 m a.s.l. Adults are often found in broad-leaved, mixed coniferous woods, rock meadows, steppes and tundras (Savchenko 1960(Savchenko , 1964Starkevich 2012). The Qinghai-Tibetan Plateau (QTP) is the largest and highest plateau in the world with an average elevation above 4000 m. The uplift and formation of the QTP has strongly infl uenced the climate, environment and evolution of biota in this region (Li & Fang 1999). The QTP has an important status in zoogeographical regionalization, which spans three biodiversity hotspots in the world, including the mountains of Southwest China, the Himalaya, and the Indo-Burma, respectively . Tibet is located in southwest China and the Qinghai-Tibet Plateau with its peculiar biodiversity in Asia. It mostly belongs to the Palaearctic Region except that southern Tibet is considered as Oriental (Su et al. 2019). The following 12 species of Vestiplex are known to occur in Tibet: Tipula (Vestiplex) adungensis Alexander, 1963, T. (V.) alyxis Alexander, 1963, T. (V.) avicularia Edwards, 1928, T. (V.) distifurca Alexander, 1942, T. (V.) eurydice Alexander, 1961, T. (V.) himalayensis Brunetti, 1911, T. (V.) nigroapicalis Brunetti, 1911, T. (V.) opilionimorpha opilionimorpha Savchenko, 1955, T. (V.) rongtoensis Alexander, 1963, T. (V.) scandens Edwards, 1928, T. (V.) subscripta Edwards, 1928 and T. (V.) zayulensis Alexander, 1963(Oosterbroek 2021. In this paper, six new species are added to the fauna of Tibet, and descriptions with illustrations of three previously known species have been added to improve the recognition of these taxa. A key to the species of Vestiplex in Tibet is presented.

Material and methods
The specimens were studied and illustrated with ZEISS Stemi 2000-c stereo microscope. Details of coloration were checked in specimens immersed in 75% ethyl alcohol (C 2 H 5 OH), to observe pruinescence specimens were examined in dry condition, some specimens were not soaked in alcohol. Genitalic preparations of males were made by immersion in a heated Lactic acid solution (C 3 H 6 O 3 > 85%, 95-97°C) for 4-6 minutes and cooled down to room temperature. After examination, genitalia were transferred to fresh glycerine (C 3 H 8 O 3 ) and stored in a microvial attached to the specimen. Type specimens are deposited in the Entomological Museum of China Agricultural University (CAU), Beijing. All pictures were adjusted and assembled into plates with Adobe Photoshop CS6. The morphological terminology mostly follows Gelhaus (2005), Cumming & Wood (2017), de Jong (2017) and Starkevich et al. (2020). Species group defi nitions follows Starkevich (2012 Alexander, 1963 12. Posterior margin of gonocoxite short thorn-like; appendages of sternite 9 strong, bifurcate at the apex; lower beak of inner gonostylus usual ..  Brunetti, 1911 Tipula (Vestiplex) deserrata group in Tibet Tipula (Vestiplex) magatama sp. nov. Tipula (Vestiplex) uncinella sp. nov.

Diagnosis
Nasus elongate, apex slightly expanded. Scape and pedicel of antenna yellow; fl agellar segments 1-4 grayish yellow, other segments pale brown, except base darker. Prescutum almost wholly reddish brown with four olive-gray striped pruinescence, median stripe broadened apically and gradually narrowing to base. Posterior margin of gonocoxite with a sharp black spine. Tergite 9 separated medially, with a pair of angular arms arising from ventral surface of tergite 9, with granulated margins, apex with a small inner projection, outer surface of arms with a pair of depressions ( Fig. 9). Outer gonostylus slender, spoon-like. Lower beak of inner gonostylus absent.

Etymology
The species is named after the shape of inner gonostylus, it looks like a 'Magatama', a curved commashaped bead from prehistoric Japan, called ' ' in China. The name is to be treated as a noun in apposition. HEAD (Figs 1, 3). Mainly brownish yellow. Occipital marking brown, elongated to vertex, broadened apically and gradually narrowing to base. Rostrum and nasus grayish yellow with brown setae, nasus elongate, apex slightly expanded. Setae on head dark brown. Antenna relatively short; scape and pedicel yellow; fl agellar segments 1-4 grayish yellow, other segments pale brown, except base darker. Proboscis and palpus brown.

Holotype
THORAX (Figs 1, 3). Mainly reddish brown, heavily grayish yellow pruinescence. Pronotum brown with yellow setae, middle black. Prescutum almost wholly reddish brown with four olive-gray striped pruinescence, median stripe broadened apically and gradually narrowing to base. Scutum brown with two olive-gray pruinose spots. Scutellum reddish brown with dark gray pruinescence. Mediotergite dark yellow with yellow pruinescence and olive-gray mid-longitudinally. Thoracic pleuron dark yellow with grayish yellow pruinescence. Anepisternum, katepisternum and anepimeron dark brown at antero-upper corners. Anatergite and katatergite yellow. Parascutellum yellow. Meron dark yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with brownish black apexes; tibiae brownish yellow with black tips. Tarsi brownish black. Setae on legs dark brown. Wing variegated brown with numerous smaller pale areas; pterostigma pale black; petiole of cell m 1 distinctly shorter than discal cell (Fig. 2). Halter with stem dark yellow, knob black.
HYPOPYGIUM . Posterior margin of gonocoxite with a sharp black spine. Tergite 9 separated medially, with a pair of angular arms arising from ventral surface of tergite 9, with granulated margins, apex with a small inner projection, outer surface arms with a pair of depressions. Sternite 9 with a median prominence. Apex of adminiculum slim. Outer gonostylus slender, spoon-like, apex circular (Fig. 5). Inner gonostylus with big concavity; inner margin black sclerotized; lower beak absent; reversed side with big concavity (Figs 6-7).

Female
Unknown or unrecognized.

Remarks
The new species is similar to T. (V.) yunnanensis Alexander, 1942 from Yunnan, China, but the latter differs in the following characters: posterior margin of gonocoxite with a slightly twisted spine; outer gonostylus long and slender; lower beak of inner gonostylus sharp.

Diagnosis
Scape and pedicel yellow, fl agellum grayish yellow. Prescutum with four grayish yellow stripes, all stripes with brown margins. Scutellum yellow with dark brown mid-longitudinal stripe. Mediotergite yellow with brown mid-longitudinal stripe. Tergite 9 separated medially, pair of angular arms arising from ventral surface of tergite 9, arms with granulated margins. Base of posterior margin of inner gonostylus without a small protrusion; upper beak slightly twisted, lower beak curved, middle concave.

Etymology
The species is named after the uncinatus of the adminiculum, the Latin adjective 'uncinella' means 'bearing hooks, barbed'.  HEAD (Figs 10,12). Mainly yellow. Occipital stripe dark gray. Rostrum and nasus yellow. Setae on head brown. Scape and pedicel yellow, fl agellum grayish yellow. Proboscis and palpus brownish yellow.

Holotype
THORAX (Figs 10,12). Mainly yellow, heavily pruinose. Pronotum brownish yellow with yellow setae. Prescutum with four grayish yellow stripes, all stripes with brown margins, median stripe broadened apically and gradually narrowing to base, lateral stripes long-oval. Scutum yellow with four grayish yellow spots, anterior margins of anterior two spots brown. Scutellum yellow with dark brown midlongitudinal stripe. Mediotergite yellow with brown mid-longitudinal stripe. Thoracic pleuron yellow. Parascutellum yellow. Meron yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with dark brown apices, tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated light brown; pterostigma brown; petiole of cell m 1 distinctly shorter than discal cell (Fig. 11). Halter with stem yellow, knob pale black with pale yellow apex.
HYPOPYGIUM 20). Gonocoxite elongated, separated from sternite 9, posterior margin with an unusually strong and slender black spine, directed chiefl y dorsally, slightly twisted. Tergite 9 separated medially, m iddle of posterior margin of lobe slightly protrude; a pair of angular black arms arising from ventral surface of tergite 9, surface of arms not smooth, with granulated margin. Sternite 9 with a median prominence. Adminiculum narrow spade-like, apex slightly furcal, ventral caudal part with a small uncinate projection. Outer gonostylus anteriorly bent, posterior margin of basal half truncated (Fig. 16). Inner gonostylus with concavity at middle, posterior margin with big semicircular bulge; base of posterior margin of inner gonostylus without a small protrusion; upper beak slightly twisted, lower beak curved, middle concave (Fig. 14). Semen pump  as shown in fi gures. Aedeagus wide, basal width about 0.5 mm, gradually narrowing towards apex, and apex not bifurcated.

Female
Unknown or unrecognized.

Remarks
The new species is similar to T. (V.) schizophallus (Alexander, 1973), but the latter differs in the following characters: arms of tergite 9 with only inner margin roughened; gonocoxite with an additional small spine (the one near posterior margin of basistylar suture); outer gonostylus with very abundant black setae from conspicuous punctures; apex of adminiculum profoundly divided into two long parallel spines, ventral caudal part without uncinatus. It is also similar T. (V.) deserrata Alexander, 1934 and T. (V.) factiosa Alexander, 1940, but the posterior margin of inner gonostylus in the latter has no protrusion. The posterior margin of tergite 9 of T. (V.) factiosa is smooth, its arms are large and strong.

Etymology
The species is named after the type locality Nayogabu Snow Mountain. HEAD (Figs 21,23). Mainly brownish yellow. Occipital mark dark brown, vertex brownish black, relatively protruded. Rostrum and nasus dark yellow, nasus relatively long. Setae on head dark brown. Scape dark yellow, pedicel pale yellow, fl agellum brownish black except fi rst fl agellar segment yellowish brown. Proboscis and palpus yellowish gray.

Holotype
THORAX (Figs 21-23). Mainly dark brown, heavily pale gray pruinescence. Pronotum brown with yellow setae. Prescutum virtually brownish black, black stripes distinct, all stripes with dark black margin. Scutum virtually brownish black. Scutellum dark yellow with brownish black mid-longitudinal stripe. Mediotergite reddish dark brown, dried with pale gray pruinescence and black mid-longitudinally. Thoracic pleuron dark brown with pale gray pruinescence. Parascutellum dark brown. Meron brown. Setae on thorax pale yellow. Legs with coxae and femora yellow, femora with brownish black apexes; tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated brown; pterostigma brown; petiole of cell m distinctly shorter than discal cell (Fig. 24). Halter with stem yellow, knob dark brown.

Female
Unknown or unrecognized.

Remarks
The new species is similar to T. (V.) zayulensis from Chayu (= Zayu) of Tibet, but the latter differs in the following characters: prescutum and scutum yellowish brown; thoracic pleuron brownish yellow; all sterno-pleurite with dark brown margin. The outer gonostylus is different from that of T. (V.) nayogabuensis sp. nov. Alexander, 1963 Figs 30-38

Diagnosis
Occipital stripe dark brown, middle wide, anterior part extended to vertex. All sterno-pleurites with dark brown margins. Tergite 9 longitudinally divided into two parts, posterior margin with long setae, and a bridge between them, a pair of angular arms arising from ventral surface of tergite 9. Inner gonostylus with concavity at middle, concavity with many bristles, lower margin of concavity with bristled raise; base of inner gonostylus inward-rolled, enwrapped black and swollen lower beak; upper beak relatively obtuse, slightly recurved. HEAD (Figs 30, 32). Mainly yellow. Occipital stripe dark brown, middle wide, anterior part extended to vertex. Rostrum and nasus yellow, nasus long. Setae on head dark brown. Scape yellow, pedicel pale yellow, fl agellum yellowish brown. Proboscis and palpus yellowish dark brown. THORAX (Figs 30, 32). Mainly brownish yellow, heavily gray pruinescence. Pronotum yellowish brown, middle and both side each with brown spot. Prescutum yellowish brown with four brown stripes, all stripes with dark brown margins. Scutum yellowish brown with four dark brown spots. Scutellum yellowish brown with dark brown mid-longitudinal stripe. Mediotergite brown with dark brown midlongitudinal stripe. Thoracic pleuron brownish yellow. All sterno-pleurite with dark brown margins. Setae on thorax yellow. Legs with coxae yellow, femora with dark brown tips; tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated pale brown; pterostigma brown; petiole of cell m 1 distinctly shorter than discal cell (Fig. 31). Halter with stem yellow, knob greyish black.
HYPOPYGIUM (Figs 33-38). Gonocoxite slightly elongated, separated from sternite 9. Tergite 9 longitudinally divided into two parts, posterior margin with long setae, and a bridge between them, a pair of angular arms arising from ventral surface of tergite 9. Sternite 9 with a pair of very short appendages, as shown in fi gure, with a long seta (Fig. 35), covered by inner gonostylus and not visible in the lateral external view. Outer gonostylus relatively small, posterior margin basally slightly wide (Fig. 34). Inner gonostylus with concavity at middle, concavity with many bristles, lower margin with bristled raise; base of inner gonostylus inward-rolled, enwrapped black and swollen lower beak; upper beak relatively obtuse, slightly recurved (Fig. 36).

Diagnosis
Scape and pedicel yellow, fi rst fl agellar segment dark yellow, other fl agellar segments bicolor, yellowish with dark brown base. Scutellum and mediotergite dark yellow. Gonocoxite elongated, separated from sternite 9, posterior margin with an unusually strong black spine, bent inward and horn-like.

Etymology
The species is named after the shape of gonocoxite, the Latin adjective 'bucera' means 'ox-horned'.  HEAD (Figs 39, 41). Mainly yellow. Occipital stripe dark brown, with brown setae. Rostrum and nasus yellow, rostrum with brown setae and nasus with dark yellow setae. Scape and pedicel yellow, fi rst fl agellar segment dark yellow, other fl agellar segments bicolor yellowish with dark brown base. Proboscis and palpus grayish yellow.

Holotype
THORAX (Figs 39,41). Mainly yellow, heavily pruinose. Pronotum yellow with yellow setae. Prescutum with four yellowish gray stripes, median stripe broadened apically and gradually narrowing to base, lateral stripes long-oval. Scutum yellow with four yellowish gray spots. Scutellum and mediotergite dark yellow. Thoracic pleuron yellow. Parascutellum pale yellow. Meron yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with dark brown apices and brown middle parts; tibiae and tarsi dark brown. Setae on legs dark brown. Wing light brown; pterostigma brown; petiole of cell m 1 and discal cell almost equilong (Fig. 40). Halter with stem brown, knob dark brown with pale tip.

Remarks
The new species is similar to T. (V.) coxitalis Alexander, 1935 from Anhui and Taiwan, China, but the latter differs in the following characters: gonocoxite ends with a slightly twisted spine; outer gonostylus narrow, fi liform; inner gonostylus without slanted bar-like raise; sternite 9 near caudal end on either side of midline with a conspicuous hair pencil of reddish bristles, these pencils not arising from basal tubercles (Men 2015). This species is similar to T. (V.) bicalcarata Savchenko, 1965 and T. (V.) maoershanensis Men & Young, 2017, but the outer gonostylus of the latter is not willow leaf shaped. The posterior margin of inner gonostylus with a fi nger-like protrusion (Men et al. 2017). Brunetti, 1911 Figs 51-57, 86 Tipula himalayensis Brunetti, 1911: 252.   HEAD (Figs 51,53). Mainly reddish yellow. Vertex with brown stripe. Rostrum and nasus dark yellow. Setae on head dark brown. Scape yellow, pedicel pale yellow, fi rst fl agellar segment yellowish brown, other fl agellar segments dark brown. Proboscis and palpus brownish yellow.

Tipula (Vestiplex) himalayensis
THORAX (Figs 51, 53). Mainly dark yellow, heavily pruinose. Pronotum dark yellow with dark yellow setae, middle pale brown and both sides with a pale brown spot. Prescutum dark yellow with four reddish brown stripes, median stripes broadened apically and gradually narrowing to base. Scutum with four reddish brown spots. Scutellum yellow with black mid-longitudinal stripe. Mediotergite yellow. Thoracic pleuron reddish yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with brownish black apices and middle portions, tibiae and tarsi brownish black. Setae on legs brownish black. Wing light brown; pterostigma pale brown; petiole of cell m 1 slightly shorter than discal cell (Fig. 52). Halter with stem yellow, knob pale yellow.

Remarks
Here, we add new distributional data and new illustrations. Yigong Township is now the current northernmost distribution of T. (V.) himalayensis. The species was considered by Brunetti to be variable but within certain limits and can be easily recognized by the pale yellow wide ring on all the femora about as far from the tip as the width of the ring. We examined other additional material, in addition to the previously indicated: 1 ♂ (paratype) Darjiling; 6000 feet a.s.l.; 22 Sep. 1908; Brunetti (USNM) / 1 ♂ (syntype) Darjiling; 20 Oct. 1905; Brunetti (BMNH(E)#246064). There is no difference in the characteristics of the femora of the three, but the wing veins are slightly different. It seems that the difference in the wing veins shown in Joseph's illustrations (Joseph 1974) can be regarded as intraspecifi c variation. But there are also some differences in the shape of inner gonostylus as shown in the illustrations, especially the beak different from others. It is necessary to make further observations on the specimens of ZSI. Brunetti, 1911 Figs 58-65

Diagnosis
Occipital stripe dark brown. Rostrum brownish yellow, nasus brown with yellow setae. Scape brownish yellow and pedicel pale yellow, fi rst fl agellar segment brownish yellow, other fl agellar segments black. Scutum yellow with four brownish black spots, which are virtually confl uent on each side, anterior two spots small; all spots with olive-gray pruinescence. Upper part of fore coxa brownish black. Inner gonostylus with upper beak sharp bearing, a semicircular striped raise at base; posterior crest produced backward. Adminiculum with a pair of long projections on both sides, longer than apex of phallic guide. HEAD (Figs 58, 60). Mainly brownish yellow. Occipital stripe dark brown. Rostrum brownish yellow, nasus brown with yellow setae. Setae on head brownish black. Scape brownish yellow, pedicel pale yellow, fi rst fl agellar segment brownish yellow, other fl agellar segments black. Proboscis and palpus black.
THORAX (Figs 58, 60). Mainly brownish yellow, heavily yellow and gray pruinescence. Pronotum brownish gray. Prescutum with four dark brownish black stripes, median stripes virtually confl uent, broadened apically and gradually narrowing to base, lateral stripes long-oval; all stripes with olive-gray pruinescence. Scutum yellow with four dark brownish black spots, which are virtually confl uent on each side, anterior two spots small, all spots with olive-gray pruinescence. Scutellum yellow with dark brown mid-longitudinal stripe. Mediotergite yellow, middlly pale gray with dark brown mid-longitudinal stripe. Scutellum and mediotergite with golden pruinescence. Thoracic pleuron with yellow pruinescence. Anepisternum and katepisternum yellow, each with a brownish black anterior lower spot. Anepimeron yellow with a small brownish black upper spot; katepimeron yellow. Meron yellow with brownish black lower part. Parascutellum yellow. Anatergite and katatergite yellow. Setae on thorax dark yellow. Legs with coxae and femora yellow, except upper part of fore coxa brownish black, femora with black apices; tibiae brownish yellow with black apices; tarsi dark black. Setae on legs black. Wing yellowish brown; pterostigma brown; petiole of cell m 1 slightly shorter than discal cell (Fig. 59). Halter with stem brown, knob grayish black with pale gray apex.

Diagnosis
Vertex with brownish black stripe. Scape, pedicel and fi rst fl agellar segment yellow, other fl agellar segments bicolor, yellowish with dark brown base. Scutellum and mediotergite dark yellow with dark gray mid-longitudinally. Prescutum dark yellow with four stripes, median stripes reddish brown, lateral stripes dark. Tergite 9 of male separated medially, with fl eshy upper tergal lobes fl attened, and two widened black sclerotized blades at a lower level.

Etymology
The species is named after the shape of blades of tergite 9, the Latin adjective 'platyphylla' refers to 'having fl at leaves'. HEAD (Figs 66,68). Mainly reddish yellow. Vertex with brownish black stripe. Rostrum and nasus dark yellow. Setae on head dark brown. Scape, pedicel and fi rst fl agellar segment yellow, other fl agellar segments bicolor, yellowish with dark brown base. Proboscis brown; palpus yellowish brown.

Holotype
THORAX (Figs 66, 68). Mainly reddish yellow with dark yellow pruinescence. Pronotum reddish yellow with yellow setae. Prescutum dark yellow with four reddish brown stripes, median stripes broadened apically and gradually narrowing to base, lateral stripes long-oval, dark reddish brown. Scutum almost entirely brown with four olive-gray pruinose spots. Scutellum and mediotergite dark yellow with dark gray mid-longitudinally. Thoracic pleuron reddish yellow. Anepisternum pale brown. Setae on thorax yellow. Legs with coxae and femora dark yellow, femora with dark brown apices and brown middle portion, tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated light brown; pterostigma brown; petiole of cell m 1 shorter than discal cell (Fig. 67). Halter with stem yellow, knob pale yellow with dark brown lower margin.

Female
Unknown or unrecognized.

Remarks
The new species is similar to T. (V.) gandharva Alexander, 1951 from India (Assam) and Myanmar, but the latter differs in the following characters: dorsolateral extensions of tergite 9 long and slender, sclerotized ventral blades much less massive; the spine of gonocoxite is smaller than this new species.

Diagnosis
Proboscis relatively short. Pronotum brownish yellow with dark yellow setae. Prescutum mainly reddish yellow with four olive-brown striped pruinescence, velvety, two middle stripes virtually fused. Gonocoxite ends hook-like, not separated from sternite 9; tergite 9 with two pairs of lobes, longitudinally divided into two parts, slightly connected, pair of emarginate arms arising from ventral surface of tergite 9. Inner gonostylus with posterior crest produced backward, base of inner gonostylus and outer basal lobe of inner gonostylus rolled inward, produced depression with row of short setulae.

Etymology
The species is named after the type locality Motuo.  HEAD (Figs 74-75). Mainly dark yellow. Vertex with dark brown stripe. Frontal tubercle relatively high. Proboscis relatively short, rostrum and nasus dark yellow. Setae on head dark brown. Scape yellow, pedicel pale yellow, fl agellum dark brown. Proboscis and palpus brownish yellow.

Material
THORAX (Figs 74-75). Mainly dark yellow, heavily pruinescence. Pronotum brownish yellow with dark yellow setae. Prescutum mainly reddish yellow with four olive-brown striped pruinescence, velvety, two middle stripes virtually fused. Scutum with four reddish brown spots, with olive-brown striped pruinescence. Scutellum and mediotergite yellow with pale brown mid-longitudinal stripes. Thorax yellow in lateral view. Pleura below anterior thoracic spiracle to middle with an irregular brownish black spot; anepisternum and katepisternum pale brownish yellow. Setae on thorax yellow. Legs with coxae and bases of femora pale yellow, apices of femora brown; tibiae and tarsi dark brown. Setae on legs brownish black. Wing light brown; pterostigma brown; petiole of cell m 1 shorter than discal cell (Fig. 76). Halter with stem dark yellow, knob brownish yellow.
HYPOPYGIUM (Figs 79-83). Gonocoxite ends hook-like, not separated from sternite 9, tergite 9 bearing two pairs of lobes, longitudinally divided into two parts, slightly connected, inner margin with dense setae; a pair of emarginate arms arising from ventral surface of tergite 9, inner margin black, other areas yellow. Sternite 9 with a median prominence, middle of sternite 9 slightly raised. Sternite 8 with abundant setae. Outer gonostylus slender and small, apex curly (Figs 79-81). Inner gonostylus, dorsal crest produced into fi shtail, base of inner gonostylus and outer basal lobe of inner gonostylus inward-rolled, produced depression with a row of short setulae, outer basal lobe of inner gonostylus large triangular (Figs 80-81).
COLORATION. Body color slightly paler than male. Antenna shorter than male, dark yellow. Legs paler than male, tibiae and tarsi dark yellow. Abdominal tergites each with three stripes, middle stripe brown, lateral stripes brownish black, abdominal 7-8 segments yellowish brown, 8th segment dark brown. Junction of each 3-9 segments brownish black (Fig. 77).

Remarks
The new species is similar to T. (V.) theowaldana Alexander, 1964 from Assam, India, but the latter differs in the following characters: basal abdominal tergites yellow; gonocoxite ends sharp and not hooked; tergite 9 with a pair of larger and elongate lobes. The species may form a new species group, perhaps together with T. (V.) theowaldana. However, we have not examined the specimens of the latter species and can therefore not say with certainty if these two species share characteristics that would separate them from existing species groups. Starkevich (2012) divided the subgenus Vestiplex into 7 species groups in his revisionary study of the global fauna. The species diversity of Vestiplex is particularly high in northern India, the Tibetan Plateau and the Hengduan Mountains, with more than 90 species currently known (Oosterbroek 2021). We added six new species to the fauna of Tibet, and placed T.