New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II

Here, we present the results of our fi eld survey in Sri Lanka and describe ten new species of Sericini: Selaserica fabriziae sp. nov., Sel. sororinitida sp. nov., Neoserica pophami sp. nov., Maladera haniel sp. nov., M. kishi sp. nov., M. windy sp. nov., M. karunaratnae sp. nov., M. hiyarensis sp. nov., M. dambullana sp. nov., and M. deenstana sp. nov. All seven of the newly described species of Maladera belong to the M. fi stulosa species group, which is an endemic radiation on the island that is characterized by entirely reduced or fused parameres. An updated key to the Maladera fi stulosa group is provided. Further, new locality records for 23 already known species are given. The genitalia and habitus of all new species are illustrated, the distribution of the new species is shown with maps.


Introduction
The fauna of the Sericini Kirby, 1837 of Sri Lanka has recently been the focus of some intensive investigations (Fabrizi & Ahrens 2014;Ranasinghe et al. 2020). Currently, the Sri Lankan fauna of Sericini comprises 81 species (Fabrizi & Ahrens 2014;Ranasinghe et al. 2020), 71 of them are endemic.
For a more thorough investigation of the fauna, we carried out a series of recent fi eld expeditions to Sri Lanka, with the aim of collecting fresh material for both morphological and molecular phylogenetic studies. While the results from the fi rst expedition in 2019 were published in Ranasinghe et al. (2020), here we present the results of the three subsequent expeditions, during which we were able to discover further ten new species belonging to the genera Maladera Mulsant & Rey, 1871, Neoserica Brenske, 1894and Selaserica Brenske, 1897. Furthermore, we report new locality records for twenty-three previously known species.  sparse and fi ne punctures, punctures glabrous; ventral margin with three strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation distinctly concave. Tarsomeres impunctate dorsally, ventrally with dense, fi ne setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is no strong longitudinal carina, fi rst metatarsomere one quarter of its length longer than dorsal tibial spur, subsequent tarsomeres lacking in holotype. Protibia moderately long, tridentate. Protarsomeres lacking in holotype.

Diagnosis
Selaserica sororinitida sp. nov. is in shape of aedeagus and external morphology similar to Sel. nitida (Candèze, 1861) and Sel. knucklensis Fabrizi & Ahrens, 2014. Selaserica sororinitida differs by the antennal club, which is longer compared to that of Sel. nitida but shorter in comparison with that of Sel. knucklensis. From both species Sel. sororinitida can also be distinguished by the shape of the aedeagus: the right paramere is in the new species shorter and less narrowed before the apex (lateral view); the phallobase is not widened distally as in Sel. knucklensis.

Etymology
The name is derived from the Latin noun 'soror' (sister) with 'nitida' from the species Sel. nitida, with reference to the strong morphological similarity with Sel. nitida (noun in nominative case).
PRONOTUM. Moderately wide, widest at base, lateral margins in basal half almost straight and weakly convergent, in anterior half weakly convex and distinctly narrowed towards anterior angles, anterior angles moderately produced and moderately sharp; anterior margin moderately produced medially, its marginal line complete; surface moderately densely and fi nely punctate, glabrous; anterior and lateral borders sparsely setose; basal margin without marginal line. Hypomeron ventrobasally not carinate. Scutellum wide, triangular, with fi ne and dense punctures, glabrous, apical portion completely impunctate.
ELYTRA. Oblong, widest in posterior third, striae distinctly impressed, fi nely and densely punctate, intervals fl at, with fi ne, sparse punctures, glabrous, only on penultimate lateral interval a few single long, erect setae; epipleural edge fi ne, ending well anterior to convex external apical angle of elytra; epipleura densely setose; apical border membranous, apex covered with a narrow rim of short microtrichomes. VENTRAL SURFACE. Dull, thorax and metacoxa with large and dense punctures, with long and dense setae; metacoxa glabrous except for a few long setae laterally; each abdominal sternite with evenly distributed fi ne and dense punctures, and with a transverse row of coarse punctures each bearing a robust seta, other punctures with a short or minute setae, penultimate sternite apically with a shiny smooth but very short chitinous border. Mesosternum between mesocoxae little narrower than maximum width of mesofemur.
LEGS. Moderately wide and shiny; femur with two longitudinal rows of setae, fi nely and moderately densely punctate; metafemur with acute anterior edge, without an adjacent serrated line, anterior longitudinal row of setae complete; posterior ventral margin almost weakly convex, weakly widened in apical half, not serrated, glabrous; posterior dorsal margin smooth, densely shortly setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.3; dorsal margin longitudinally convex, with two groups of spines, basal one at fi rst quarter, apical one at three quarters of metatibial length, without a serrated line; lateral face longitudinally convex, with very sparse and fi ne punctures, glabrous; ventral margin serrated, with fi ve robust equidistant spines; medial face smooth; apex interiorly near tarsal articulation concavely emarginate. Tarsomeres dorsally impunctate and glabrous, ventrally with dense, long and robust setae; metatarsomeres ventrally with a serrated ridge, beside it without additional strong longitudinal carina; fi rst metatarsomere one third of its length longer than dorsal tibial spur, distinctly shorter than following two tarsomeres combined. Protibia moderately long, tridentate, basal tooth smaller than the two distal ones. Protarsomeres ventrally with long and dense yellow setae forming a setose pad, protarsal claws symmetrical, basal tooth of claws sharply pointed.

Diagnosis
Neoserica pophami sp. nov. differs from the very similar N. kalaarensis Fabrizi & Ahrens, 2014 in shape of parameres: the right paramere is slightly longer, medially distinctly bent (lateral view) and much wider (dorsal view).

Etymology
The new species is named after Mr Sam Popham, founder of the NIFS Arboretum (noun in the genitive case).

Type material
Holotype SRI LANKA • ♂; "X-SR0346, Sri Lanka,Matale District,Dambulla,NIFS Arboretum,7.85783ºN,80.67391ºE,167m, HEAD. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and convergent anteriorly, anterior angles moderately rounded, anterior margin slightly sinuated medially, all margins moderately refl exed, lateral margins produce an indistinct angle with the ocular canthus; surface weakly convex medially, densely punctate, coarse and fi ne punctures mixed, with numerous erect setae; frontoclypeal suture indistinctly incised, not elevated and weakly angled medially; smooth area anterior to eye fl at, three times as wide as long; ocular canthus moderately long and wide (one third of ocular diameter), fi nely and densely punctate, with a single terminal seta. Frons with fi ne, sparse punctures, with a few erect setae beside eyes. Eyes moderately large, ratio diameter/interocular width: 0.75. Antenna with ten antennomeres; club with fi ve antennomeres and straight, as long as the remaining antennomeres combined. Mentum elevated and slightly fl attened anteriorly.
PRONOTUM. Moderately transverse, widest at base, lateral margins in basal half almost straight and moderately convergent anteriorly, in anterior half weakly convex and evenly convergent anteriorly, anterior angles distinctly produced and moderately acute, posterior angles blunt; anterior margin almost not produced medially, with a robust and complete marginal line, base of pronotum without marginal line; surface fi nely and densely punctate, punctures glabrous; lateral and lateral anterior margin sparsely setose; hypomeron carinate, not ventrally produced. Scutellum wide, triangular, at apex moderately pointed, with fi ne, moderately dense punctures, glabrous.
ELYTRA. Widest shortly behind the middle, striae weakly impressed, fi nely and moderately densely punctate, intervals fl at, with fi ne and evenly moderately dense punctures, except a few short setae on lateral intervals glabrous; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra with a fi ne rim of microtrichomes (100 ×). VENTRAL SURFACE. In major part dull, some parts shiny, coarsely and densely punctate, metasternum sparsely covered with fi ne, short or very minute setae, metacoxa glabrous, with a few long setae laterally; abdominal sternites fi nely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta, the row of punctures on the fi rst visible basal sternite fused to a robust transverse carina bearing the setae, before the carina the fi ne punctation extremely dense but glabrous. Mesosternum between mesocoxae 1.5 times as wide as the width of mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.65. Pygidium strongly convex and shiny, fi nely and moderately densely punctate, without smooth midline, glabrous except some longer setae along the apical margin.
LEGS. Wide and moderately long; femur with two longitudinal row of setae, fi nely and sparsely punctate; metafemur shiny, behind the posterior longitudinal row of setae punctures fi ner and slightly denser, anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, strongly widened, posterior margin fi nely serrated over its entire length dorsally, with just a few short setae basally. Metatibia wide and fl attened, short, widest at apical second third of metatibial length, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group at fi rst third, apical group at two thirds of metatibial length, basally with a few short robust single spines, on basal quarter beside dorsal margin with a short serrated line; lateral face weakly longitudinally convex, fi nely, superfi cially and sparsely punctate, glabrous, widely smooth along the middle; ventral margin fi nely serrated, with four robust equidistant setae; medial face smooth and glabrous; apex fi nely serrated, interiorly near tarsal articulation weakly concavely sinuate. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, ventrally robustly densely setose; metatarsomeres with a strongly serrated ridge and a smooth subventral longitudinal carina; fi rst metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. AEDEAGUS. Fig. 3I-K.

Female
Length: 6.8-7.2 mm, length of elytra: 4.6-5.0 mm, width: 3.7-4.0 mm. Eyes as large as in male; antennal club shorter little than remaining antennomeres combined, the basal joint of club equals only ⅓ of the length of the club; pygidium weakly convex.

Etymology
The new species is named for Suresh Benjamin's son, Haniel P. Benjamin (noun in apposition).

Type material
Holotype  HEAD. Labroclypeus short and trapezoidal, wider than long, widest at base, lateral margins strongly convex and convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly refl exed, anterior margin almost weakly sinuate medially; surface slightly convex, fi nely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture indistinctly incised and bluntly bent medially; smooth area in front of eye approximately three times as wide as long; ocular canthus long and narrow, minutely and superfi cially punctate, with a single short terminal seta. Frons with fi ne, dense punctures, with a few long erect setae in larger punctures, setae on disc less dense. Eyes large, ratio of diameter/interocular width: 0.79. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Wide, widest at base, lateral margins in basal half straight and weakly convergent, in anterior half weakly convex and narrowed to anterior angles, anterior angles moderately produced and sharp, anterior marginal line fi ne and complete, anterior margin convexly produced medially; surface fi nely densely punctate, with moderately dense, short and fi ne setae and dense long erect setae being directed anteriorly; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures, with short, dense, fi ne setae.
ELYTRA. Short oval, widest shortly behind middle, striae distinctly impressed, fi nely and densely punctate, intervals weakly convex, with moderately fi ne, dense punctures and with dense, fi ne, short setae as well as with sparse long erect setae; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Shiny, thorax and metacoxa with large and dense punctures, sparsely setose, metacoxa with minute adjacent setae in the punctures except for numerous long setae laterally, apical margin weakly convex, without a wide rim of long white microtrichomes; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, 3 rd sternite with a sharp median tubercle being half as high as sternite length, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/1.93. Pygidium moderately convex, moderately fi nely and densely punctate, without smooth midline, punctures with short and dense, adjacent setae as well as with moderately dense, long, erect setae. LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half, fi nely serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly only very little narrowed, ratio width/ length: 1/2.5, dorsally sharply carinate, with two groups of spines, basal shortly behind middle, apical one at three quarters of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing a single robust spine and beside them a longitudinal serrated line; lateral face almost fl at, with dense, large punctures and with minute setae in the punctures; ventral margin with fi ve strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly concave. Meso-and metatarsomeres fi nely and sparsely punctate but glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; fi rst metatarsomere as long as following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.

Female
Unknown.

Diagnosis
Maladera kishi sp. nov. is in external morphology very similar to M. haniel sp. nov. and differs by the lack of a tubercle on abdominal sternite III as well as by the shape of parameres: the fused parameres (distal aedeagal process) is simply pointed and not bifurcate, while the lateral process on the left side is more robust.

Etymology
The new species is named for Suresh Benjamin's daughter, Kishi P.L. Benjamin (noun in apposition).

Type material
Holotype SRI LANKA • ♂; "X-SR0724, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X- HEAD. Labroclypeus short and trapezoidal, wider than long, widest at base, lateral margins strongly convex and convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly refl exed, anterior margin not sinuate medially; surface slightly convex, fi nely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture indistinctly incised and bluntly bent medially; smooth area in front of eye approximately three times as wide as long; ocular canthus long and narrow, minutely and superfi cially punctate, without a short terminal seta. Frons with fi ne, dense punctures, with a few long erect setae in larger punctures. Eyes large, ratio of diameter/interocular width: 0.81. Antenna yellow, with ten antennomeres; club with three antennomeres, little longer than remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Wide, widest at base, lateral margins in basal half straight and weakly convergent, in anterior half weakly convex and narrowed to anterior angles, anterior angles moderately produced and sharp, anterior marginal line fi ne and complete, anterior margin convexly produced medially; surface fi nely densely punctate, with moderately dense, short and fi ne setae and dense long erect setae being directed anteriorly; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures, with short, dense, fi ne setae.
ELYTRA. Short oval, widest shortly behind middle, striae distinctly impressed, fi nely and densely punctate, intervals weakly convex, with moderately fi ne, dense punctures and with dense, fi ne, short setae as well as with sparse long erect setae; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Shiny, thorax and metacoxa with large and dense punctures, sparsely setose, metacoxa with minute adjacent setae in the punctures except for numerous long setae laterally, apical margin weakly convex, without a wide rim of long white microtrichomes; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/1.8. Pygidium moderately convex, moderately fi nely and densely punctate, without smooth midline, punctures with short and dense, adjacent setae as well as with dense, long, erect setae.
LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half, fi nely indistinctly serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly only very slightly narrowed, ratio width/length: 1/2.4, dorsally sharply carinate, with two groups of spines, basal well behind middle, apical one at four fi fths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing a single robust spine and beside them a longitudinal serrated line; lateral face almost fl at, with dense, large punctures and with fi ne setae in the punctures; ventral margin with fi ve strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly concave. Meso-and metatarsomeres fi nely and sparsely punctate but glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; fi rst metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.

Diagnosis
The new species is very similar to M. pubescens (Arrow, 1916) and M. dambullana sp. nov. as well as M. bisornata Fabrizi & Ahrens, 2014. Maladera windy sp. nov. differs from the fi rst two by the distal aedeagal lobe (i.e., the fused parameres) having a basal lateral lobe with grater-like surface, the ventral process of phallobase is shorter, less pointed and not mesally extended; from M. bisornata, M. windy sp. nov. differs by the longer and slightly refl exed distal aedeagal lobe (i.e., the fused parameres).

Etymology
The name of the new species is derived from 'Windy Holiday Bungalow', where the research group stayed during the second expedition in Knuckles region (noun in apposition).

Type material
Holotype SRI LANKA • ♂; "X-SR0769, Sri Lanka,Deenston,Knuckles South,7.35771ºN,80.85006ºE,980m, HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins moderately refl exed, anteriorly distinctly sinuate medially; surface fl at, fi nely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture fi ne and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, fi nely and densely punctate, with a single short terminal seta. Frons with fi ne, dense punctures, with dense short and sparse long erect setae. Eyes large, ratio of diameter/interocular width: 0.85. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Moderately wide, widest at base, lateral margins weakly convex and evenly narrowed anteriorly, anterior angles strongly produced and sharp, anterior marginal line fi ne and complete, anterior margin weakly produced medially; surface fi nely and densely punctate, with dense moderately long setae being bent posteriorly on entire disc and with a few sparse longer setae being directed anteriorly; anterior and lateral borders setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures, with fi ne and dense adjacent setae.
ELYTRA. Short oval, widest shortly behind middle, striae indistinctly impressed, fi nely and densely punctate, intervals fl at, with fi ne, very dense punctures, with numerous fi ne setae similar to those of the pronotum and a few sparser ones being longer and erect or directed anteriorly, in particular on lateral intervals; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely setose, metacoxa glabrous except for numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/1.9. Pygidium moderately convex, fi nely and very densely punctate, without smooth midline, punctures with short and moderately dense, adjacent setae, along the apical margin with a few long erect setae.
LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half and very indistinctly serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.6, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fi fths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and beside them a longitudinal serrated line; lateral face almost fl at, with dense, fi ne punctures and with minute setae; ventral margin with fi ve strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso-and metatarsomeres fi nely and densely punctate and setose dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; fi rst metatarsomere little shorter than following two tarsomeres combined and a little longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.

Etymology
The new species is named after Prof. Inoka Karunaratne (University of Peradeniya), in gratitude for her kind support for this project (noun in the genitive case).  (Fig. 5D) AND COLOURATION. Body oval, brown, antenna, ventral side, and legs yellowish, dorsal surface with iridescent shine, densely shortly setose, elytra with numerous single erect setae.
HEAD. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex and convergent to widely rounded anterior angles, lateral border and ocular canthus producing an blunt angle, margins moderately refl exed; anterior margin weakly emarginate medially; surface shiny, fl at, fi nely and coarsely, densely punctate, with a few erect setae in larger punctures and minute setae in the remaining punctures; frontoclypeal suture fi nely incised and weakly curved; smooth area in front of eye three times as wide as long; ocular canthus long and narrow, sparsely fi nely punctate, with a single short terminal seta. Frons with toment and iridescent shine, with fi ne, dense punctures and short, erect setae in punctures, with a few long, erect setae on disc and beside eyes. Eyes extremely large, ratio of diameter/ interocular width: 1.1. Antenna yellow, with ten antennomeres; club with three antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Narrow, widest at base, lateral margins weakly convex and convergent anteriorly; anterior angles moderately produced and sharp; anterior marginal line fi ne and complete, anterior margin weakly produced medially; surface fi nely and densely punctate, with short erect setae in punctures, and a numerous longer erect setae in anterior part; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures and setae as in pronotum.
ELYTRA. Short oval, widest at middle, striae indistinctly impressed, fi nely and densely punctate, intervals fl at, with fi ne, dense punctures, with short setae in punctures; odd intervals with numerous longer, erect setae around which the smaller setae are lacking circularly; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border membranous, apex covered with a rim of short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely shortly setose; metacoxa with numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a narrow, shiny smooth chitinous border. Mesosternum between mesocoxae as wide as mesofemur, with a semicircular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/2.28. Pygidium weakly convex, fi nely densely punctate, with short dense setae, and numerous long setae on apical half.
LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose; posterior ventral margin straight, strongly widened in apical half and very indistinctly serrate apically; posterior dorsal margin not serrated, densely setose. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.4, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fi fths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and a longitudinal serrated line in basal half; lateral face longitudinal convex, with dense, fi ne punctures and with fi ne white setae in punctures; ventral margin fi nely serrate, with fi ve strong spines equidistant from each other; medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso-and metatarsomeres dorsally densely and fi nely punctate, and densely setose, ventrally with robust, dense, short setae; metatarsomeres with a strongly serrated ventral ridge, with a strong longitudinal carina beside it; fi rst metatarsomere as long as two following tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth, protarsal claws asymmetric, basal tooth of inner claw widened and bluntly truncate at apex. AEDEAGUS. Fig. 5A-C.

Etymology
The new species is named after its type locality 'Hiyare' (adjective in the nominative singular).

Description
MEASUREMENTS. Length: 6.8 mm, length of elytra: 4.8 mm, width: 4.1 mm. HABITUS (Fig. 5H) AND COLOURATION. Body oval, reddish brown, frons, disc of pronotum and a few spots on elytra dark brown, antenna and legs yellow, dorsal surface with iridescent shine, densely and shortly setose, elytra with numerous single erect setae.
HEAD. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins moderately refl exed; anterior margin very weakly emarginate medially; surface shiny, weakly elevated medially, fi nely and densely punctate, distance between punctures equal their diameter, with a few erect setae in larger punctures; frontoclypeal suture fi ne and weakly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus moderately long and narrow, impunctate, with a single short terminal seta. Frons with toment and iridescent shine, with fi ne, dense punctures and minute setae in punctures, with a few long, erect setae on disc and beside eyes. Eyes large, ratio of diameter/interocular width: 0.79. Antenna yellow, with ten antennomeres; club with three antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Moderately wide, widest at base, lateral margins in basal half straight and convergent, in anterior half weakly convex and convergent anteriorly; anterior angles moderately produced and sharp; anterior marginal line fi ne and complete, anterior margin weakly produced medially; surface fi nely and densely punctate, with short setae in punctures, otherwise glabrous; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures, along midline narrowly impunctate. ELYTRA. Short oval, widest at posterior third, striae indistinctly impressed, fi nely and densely punctate, intervals fl at, with fi ne, dense punctures, with short setae in punctures; odd intervals with a few impunctate dots which are darker and each bear at centre an erect seta; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border membranous, apex covered with a rim of short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely shortly setose; metacoxa with numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a narrow, shiny smooth chitinous border. Mesosternum between mesocoxae as wide as mesofemur, with a semicircular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/2.24. Pygidium weakly convex, fi nely densely punctate, with short dense setae, and numerous long setae along apical margin.
LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose; posterior ventral margin straight, strongly widened in apical half and very indistinctly serrate apically; posterior dorsal margin not serrated, densely setose. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.4, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fi fths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and beside them a longitudinal serrated line; lateral face longitudinal convex, with dense, fi ne punctures and with short setae in punctures; ventral margin fi nely serrate, with fi ve strong spines equidistant from each other; medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso-and metatarsomeres dorsally sparsely and very fi nely punctate and with minute setae in punctures, ventrally with robust, dense, short setae; metatarsomeres with a strongly serrated ventral ridge, beside it with a strong longitudinal carina; fi rst metatarsomere as long as two following tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth, distal protarsomeres lacking in holotype.

Female
Unknown.

Diagnosis
The new species is very similar to M. pubescens (Arrow, 1916). Maladera dambullana sp. nov. differs from M. pubescens by the shape of the aedeagus: the ventral distal lobe (i.e., the fused parameres) is not extended mesally but constant in width over its entire length.

Etymology
The name of the new species is derived from its type locality 'Dambulla' (adjective in nominative case singular).

Type material
Holotype SRI LANKA • ♂; "X-SR0269, Sri Lanka,Matale District,Dambulla,NIFS Arboretum,7.85766ºN,80.67474ºE,174m, HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly refl exed, anteriorly shallowly sinuate medially; surface slightly convex, fi nely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture almost invisible and strongly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, minutely and superfi cially punctate, with a single short terminal seta. Frons with fi ne, dense punctures, with long erect setae in the punctures. Eyes large, ratio of diameter/interocular width: 0.85. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Moderately wide, widest at base, lateral margins weakly convex and evenly narrowed to the anterior third, anteriorly stronger convex, anterior angles strongly produced and sharp, anterior marginal line fi ne but complete medially, anterior margin weakly produced medially; surface fi nely and densely punctate, with dense moderately long setae being bent posteriorly on entire disc and with a few sparse longer setae being directed anteriorly; anterior and lateral borders setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures, with fi ne and dense adjacent setae.
ELYTRA. Short oval, widest shortly behind middle, striae indistinctly impressed, fi nely and densely punctate, intervals fl at, with fi ne, very dense punctures, with numerous fi ne setae similar to those of the pronotum and a few sparser ones being longer and erect or directed anteriorly, in particular on lateral intervals; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely setose, metacoxa glabrous except for numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/1.93. Pygidium moderately convex, fi nely and very densely punctate, without smooth midline, punctures with short and moderately dense, adjacent setae, along the apical margin with a few long erect setae.
LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half and very indistinctly serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.18, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and beside them a longitudinal serrated line; lateral face almost fl at, with dense, fi ne punctures and with minute setae in the punctures; ventral margin with fi ve strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso-and metatarsomeres fi nely and densely punctate and setose dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; fi rst metatarsomere as long as following two tarsomeres combined and a little longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.

Diagnosis
Maladera deenstana sp. nov. is in external shape and shape of aedeagus similar to M. weligamana (Brenske, 1898). The new species differs from M. weligamana by the mesally more compressed aedeagus and having the distal part less narrowed towards apex (all lateral view); the apical part is less narrowed a quarter before apex than in M. weligamana (dorsal view).

Etymology
The new species is named after its type locality 'Deenston' (adjective in the nominative singular).
HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins moderately refl exed, anteriorly distinctly emarginate medially; surface shiny, convexly elevated medially, fi nely and densely punctate, distance between punctures equal their diameter, with a few erect setae; frontoclypeal suture fi ne and weakly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, fi nely and densely punctate, with a single short terminal seta. Frons dull, with fi ne, irregularly sparse punctures, with a few longer, adpressed setae on disc and beside eyes. Eyes moderately large, ratio of diameter/interocular width: 0.66. Antenna yellow, with ten antennomeres; club with three antennomeres, little shorter than remaining antennomeres combined. Mentum elevated and anteriorly fl attened.
PRONOTUM. Moderately wide, widest at base, lateral margins weakly convex and evenly narrowed anteriorly, anterior angles moderately produced and moderately sharp, anterior marginal line fi ne and complete, anterior margin weakly produced medially; surface fi nely and densely punctate, with minute  setae in punctures, otherwise glabrous; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fi ne and dense punctures, along midline impunctate.
ELYTRA. Short oval, widest at posterior third, striae indistinctly impressed, fi nely and densely punctate, intervals fl at, with fi ne, dense punctures, with minute setae in punctures and a few moderately dense, short setae on lateral intervals and posterior half; epipleural edge fi ne, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border membranous, apex covered with a rim of short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely minutely setose, metacoxa with numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fi ne and dense punctures bearing each a fi ne seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a narrow, shiny smooth chitinous border. Mesosternum between mesocoxae as wide as mesofemur, with a semicircular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/2.0. Pygidium lacking in holotype.
LEGS. Short and wide; femur with two longitudinal rows of setae, fi nely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose; posterior ventral margin straight, strongly widened in apical half and very indistinctly serrate apically; posterior dorsal margin not serrated, densely setose. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.9, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fi fths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines; lateral face longitudinal convex, with sparse, fi ne punctures and with minute setae in punctures; ventral margin fi nely serrate, with fi ve strong spines equidistant from each other; medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso-and metatarsomeres impunctate and glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally glabrous, with a strongly serrated ridge, beside which is a strong longitudinal carina; fi rst metatarsomere as long as two following tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth, distal protarsomeres lacking in holotype.

Female
Unknown.

Distribution
See Fig. 7J.  12. Aedeagus strongly compressed at middle, its distal part strongly narrowed towards apex (lateral view) and distinctly narrowed a quarter before apex (dorsal view   Fabrizi & Ahrens, 2014