Three new species of Anacharis Dalman, 1823 (Hymenoptera: Figitidae), with revised taxonomy and distribution records of Palaearctic and Indomalayan species

Abstract. An update of the current knowledge of Anacharis Dalman, 1823 for the Palaearctic and Indomalayan regions is given. The previously known Palaearctic species Anacharis antennata Belizin, 1951, Anacharis eucharoides (Dalman, 1818), Anacharis immunis Walker, 1835 and Anacharis parapsidalis Belizin, 1951 are redescribed. Three new species are described: Anacharis fergussoni sp. nov. from Europe, Anacharis norvegica sp. nov. from Norway and Anacharis belizini sp. nov. from Thailand, the first recorded Indomalayan species for the genus. Anacharis gracilipes Ionescu, 1969, is synonymized with A. eucharoides, while Anacharis flavidicornis Kieffer, 1910 is transferred to the genus Aegilips Haliday, 1835, resulting in Aegilips flavidicornis (Kieffer, 1910) comb. nov. Diagnostic characters and data about the biology, distribution and affinities with other species of Anacharis are discussed. An identification key for the Palaearctic and Indomalayan species of Anacharis is given.

Anacharis is the second most diverse of the Anacharitinae genera after Aegilips, and includes 21 described species.Anacharis does not have a scutellar spine, a trait shared with Aegilips, Calofigites, Hexacharis, Proanacharis and Solenofigites.Among them, Aegilips and Hexacharis are the closer ones to Anacharis (Buffington et al. 2007(Buffington et al. , 2012)), but can be distinguished from them by an elongated petiole (at least as long as metacoxa, usually longer) with smooth surface (restrepo-ortiz & Pujade-villar 2010).
Anacharis flavidicornis was described by Kieffer (1910) from Central Asia.The knowledge of Eastern Palaearctic Anacharis was extended by Belizin (1951), who cited A. immunis as occurring in the russian Far East and described two new species: Anacharis antennata from Central Asia and A. parapsidalis from the russian Far East.Belizin (1961) also cited A. eucharoides for the westernmost parts of Central Asia (Chelyabinsk oblast in russia).The Western portion of the Palaearctic region had an scarcer record of Anacharis, with only three species having been collected: Anacharis eucharoides and A. immunis cited from Eastern and Central Europe, and A. gracilipes, exclusively present in romania.
In this study, we describe A. belizini sp.nov., A. fergussoni sp.nov., and A. norvegica sp.nov., the first one being the first Indomalayan species of the genus.The Palaearctic species A. antennata, A. eucharoides, A. immunis and A. parapsidalis are redescribed and their known distribution areas are extended, while A. flavidicornis is moved to Aegilips and A. gracilipes is described as a junior synonym of A. eucharoides.Diagnostic characters for these species are given, and data about morphological features, distribution area and biology are discussed.

Material and methods
For this study 137 undetermined specimens were examined: 57 males and 80 females.Morphological terms used are those of Richards (1977), Ronquist (1995) and Ros-Farré et al. (2003).All measurements are relative except for the body length.The antennal formula includes scape, pedicel and flagellomeres length and relative width in brackets.
The undetermined specimens examined in this study are from the NHM, CNC, MNHN and uLg (see below for abbreviations).The type material studied belongs to Anacharis ensifera, A. flavidicornis, A. gracilipes, A. immunis, A. tincta and A. typica and is deposited in the NHM, MGAB and ZMB; additional material of A. eucharoides comes from Lu and is deposited in CNC, Lu, MNHN and uB.
The images included were made in the 'Serveis Científico-Tècnics' of the University of Barcelona.The field-emission gun environmental scanning electron microscope (FEI Quanta 200 ESEM) was used for high-resolution imaging, under a low voltage (12.0 kv) and without gold-coating of the specimens in order to preserve the material.

Diagnosis
Species very similar to A. eucharoides, from which it can be distinguished by always having a completely smooth mesoscutum and scutellum, strongly carinate notauli, with a short median scutellar carina at scutellar apex (mesoscutum and notauli weakly carinate in some specimens of A. eucharoides while smooth in others, the scutellum is alutaceous and lacks median carina).

Type locality
Placodeal sensilla starting at F1 and abundant in all flagellomeres in both sexes.
MesosoMa.Pronotum smooth and pubescent, with some oblique carinae in ventral region (Fig. 1B).Mesoscutal width 1.1 times its length in dorsal view.Mesoscutum smooth and shiny; almost glabrous except for a few setae.Notauli complete with transverse carinate sculpture; median mesoscutal furrow short and almost not present (Fig. 1A).Lateral region of mesoscutum smooth; parapsidal signum and parascutal sulcus absent, with a line of dense pubescence instead.Scutellar length 0.7 times that of mesoscutum in dorsal view.Scutellum smooth and shiny (Fig. 1A).

Biology
unknown.

Diagnosis
Species morphologically very similar to A. antennata, except for some weak transverse carinae at the edges of the notauli, alutaceous scutellum and a weak posterior carina on the scutellar foveae (in A. antennata, mesoscutum and scutellum smooth, scutellar foveae defined by a strong posterior carina).
CoLoration.Head, mesosoma and metasoma black.Mandibles yellowish brown with darker teeth.Antennae dark yellowish brown.Legs yellowish brown with darker coxae.veins of wings dark brown.
head.Triangular-shaped in anterior view.Face smooth, covered with sparce white setae.Width of head 2 times its length in dorsal view and 1.3 times its height in anterior view.Malar sulcus coriaceous, 0.6 times height of compound eye.Transfacial line same length as height of compound eye.Diameter of toruli larger than inter-toruli distance and torulus to compound eye distance.Clypeus smooth, glabrous, shortly convex.occipital and postocular carinae absents.Compound eyes glabrous.In both sexes PoL:ooL:LoL ratio = 7:4:4, ocelli diameter being 2.5.Frons, gena and occiput smooth, shiny and glabrous except for few sparse setae.
MesosoMa.Pronotum smooth, shiny, with some lower transverse carinae, covered by short white setae (Fig. 2D).Mesoscutal width 1.2 times its length in dorsal view.Mesoscutum smooth and shiny, almost glabrous except for a few setae; some weak transverse carinae at edges of notauli, extended in anterior region between notauli in some specimens (Fig. 2A-B).Notauli complete and clearly excavated; transverse internal carinae ranging from being absent (Fig. 2A) to present (Fig. 2B); median mesoscutal furrow absent.Lateral region of mesoscutum smooth except for a few isolated punctures; parapsidal signum and parascutal sulcus, absent.Scutellar length 0.8 times that of mesoscutum in dorsal view.Scutellum alutaceous, never areolate (Fig. 2A-B).Scutellar foveae rounded, large, alutaceous, with short oblique internal ridges in some specimens, basally defined by weak carina (Fig. 2A-B); lateral pits of scutellar foveae absent.Interfoveal line present.Circumscutellar carina complete, clearly defined, not projected at scutellar apex.Mesopleuron smooth, glabrous, shiny, with transverse groove not internally carinate; some anterior oblique carinae present.Mesopleural triangle smooth, densely pubescent.Metanotal troughs alutaceous, glabrous.Propodeum coriaceous, pubescent, divided into large cells; central area with one large upper cell partially divided by median carina and four lesser smaller cells.
Wings.Pubescent.radial cell of forewing closed, 2.6 times as long as wide.Marginal pubescence of forewing denser at apical third.
MetasoMa.Longer than head + mesosoma.Petiole longer than metacoxa, smooth, shiny, weak dorsal carinae present in last third of the petiole.Third metasomal tergum 2.5 times as long as fourth tergum in dorsal view.Fifth, sixth, seventh metasomal tergae visible in dorsal view.Metasomal terga smooth, glabrous, punctate in anterior region of each tergum, more distinct from T4 to T7.

Taxonomic remarks
The holotype of A. eucharoides is lost (Fergusson 1986); we studied the lectotypes A. tincta and A. typica designated by Fergusson (1986) instead.regarding A. gracilipes syn.nov., in the original work of Ionescu, the different coloration and longer radial cell and petiole are cited as differences for establishing a new species.After examining the type material of A. gracilipes as well as the lectotypes of A. eucharoides designated by Fergusson and series of undetermined material, we concluded that the coloration alone cannot be taken into account when distinguishing between species, and the morphometric differences cited by Ionescu (1969) fall within the ranges of intraspecific variability.

Diagnosis
Species easily distinguishable from most Eurasian species of Anacharis by the weakly excavated notauli which tend to disappear in its anterior region in some individuals (in A. antennata, A. eucharoides, A. parapsidalis, A. belizini sp.nov.and A. fergussoni sp.nov., notauli always complete, deeply excavated, internally carinate).This character is also shared with A. norvegica sp.nov., but A. immunis can be distinguished by always having a smooth mesoscutum and big cells in the scutellum (mesoscutum carinate in its anterior region and scutellum densely covered by small cells in A. norvegica sp.nov.).
MetasoMa.Shorter than head + mesosoma.Petiole as long as metacoxa, smooth and shiny.Third metasomal tergum 2.8 times longer than fourth tergum in dorsal view.Fifth, sixth, seventh metasomal terga visible in dorsal view.Metasomal terga smooth and glabrous, punctate in anterior region of each tergum, more distinct from T4 to T7.

Taxonomic remarks
Anacharis immunis and A. ensifera were described as separate species by Walker (1835).In Fergusson (1986) they were synonymized.After examining the type material and series of undetermined material, we conclude that A. immunis is a valid species with a high variability in the scutellar sculpture (from completely smooth to softly areolate) and in the scutellar foveae (clearly defined by a carina in some specimens while in others there is a coarse band at the base of the foveae instead of the basal carina).The holotype of A. immunis has a smooth scutellum and scutellar foveae without basal carina, while the lectotype of A. ensifera has an areolate scutellum and a basal carina in the scutellar foveae; other specimens reflect intermediate states of those characters.The other synonymy established by Fergusson was A. rufiventris, but we could not examine the type material.Fergusson (1986) mentioned the petiole of females being shorter than the metacoxa, while it is as long as the metacoxa in males.After examining the type material and other specimens we did not see the differences mentioned by Fergusson; both sexes present the petiole as long as the metacoxa.

Diagnosis
Species easily distinguishable from other Palaearctic Anacharis species by the strongly areolate scutellum and coarsely sculptured pronotum (pronotum smooth with some weak basal carinae and scutellum smooth to alutaceous, sometimes presenting weak carinae at the margins but never strongly areolate in A. antennata, A. eucharoides and A. immunis).
Wings.Pubescent.radial cell of forewing closed, 2.4 times as long as wide.Marginal pubescence of forewing denser at the apical third.
MetasoMa.Longer than head + mesosoma.Petiole about 2.0 times as long as metacoxa, smooth and shiny.Third metasomal tergum 1.8 times as long as fourth tergum in dorsal view.Fifth, sixth, seventh metasomal terga visible in dorsal view.Metasomal terga smooth and glabrous, not punctate.

Biology
unknown.

Distribution
Palaearctic.Known from Russia (Belizin 1951); first citation for Romania and Japan.

Comments
According to our sources, the type material of A. antennata was deposited in the Zoological Institute of St. Petersburg (ZIN), but is now lost.The specimens of our study were determined with the help of Belizin's original descriptions.

Diagnosis
Species similar to A. antennata, from which it can be distinguished by having parallel oblique ridges covering most of the pronotal surface (pronotum carinate sculpture reduced only to a few short ridges at the lower region in A. antennata).

Etymology
The specific name was chosen to honor the Russian entomologist V.I.Belizin, whose work unveiled much of the current knowledge of the Asian Anacharitinae.
CoLoration.Head, mesosoma and metasoma black.Mandibles yellowish brown with darker teeth.Antennae brown with darker first segment.Legs yellowish brown with darker coxae, third coxa black.veins of wings brownish.
head.Triangular-shaped in anterior view.Face smooth, covered with abundant white setae (Fig. 3A).Width of head 1.3 times its height in front view and 1.8 times its length in dorsal view.Malar sulcus coriaceous, 0.8 times height of compound eye.Transfacial line length equal to compound eye height.Diameter of the toruli larger than inter-toruli distance and torulus to compound eye distance.Clypeus shortly defined, convex, densely covered by pubescence.Occipital and postocular carinae absent.Compound eyes glabrous.In females, PoL:ooL:LoL ratio = 7.5:5:3, being ocelli diameter 2.5 in females.Frons, gena and occiput smooth and densely pubescent.
Wings.Pubescent.radial cell of for ewing closed, 2.8 times as long as wide.Marginal pubescence of for ewing denser at the apical third.
MetasoMa.Longer than head + mesosoma.Petiole about 2.0 times as long as metacoxa, smooth and shiny.Third metanotal tergum 1.5 times as long as fourth tergum in dorsal view.Fifth, sixth, seventh metanotal terga visible in dorsal view.Metanotal terga smooth and glabrous, punctate in anterior region of each tergum, more distinct from T4 to T7.

Biology
unknown.

Distribution
Indomalayan.Known only from Thailand.

Diagnosis
Species very similar to A. parapsidalis, from which can be distinguished by having smooth pronotum with only some short carinae in its lower region (pronotum strongly carinate in all its surface in A. parapsidalis).It is also very similar to the Nearctic species A. melanoneura, except for having alutaceous and more strongly areolate scutellum, a complete median carina in the propodeum and placodeal sensilla starting at F1 (in A. melanoneura, the scutellum is smooth and not so strongly areolate, the median propodeal carina is incomplete and placodeal sensilla start at F2). head.Triangular-shaped in anterior view.Face smooth, covered with abundant white setae.Width of head 1.3 times its height in anterior view and 2.4 times its length in dorsal view.Malar sulcus coriaceous, 0.7 times height of compound eye.Transfacial line length 1.1 times height of compound eye.Diameter of the toruli shorter than inter-toruli distance, but bigger than torulus to compound eye distance.Clypeus shortly defined, convex, densely covered by facial pubescence.Occipital and postocular carinae absent.Compound eyes glabrous.In both sexes PoL:ooL:LoL ratio = 7:5:3, ocelli diameter being 3. Frons and occiput smooth covered by some scarce hyaline hairs, gena densely pubescent.
MesosoMa.Pronotum alutaceous, densely pubescent, slightly coarse in most of its surface, presence of some short carinae in its ventral region (Fig. 4B).Mesoscutal width 1.2 times its length in dorsal view.Mesoscutum smooth, shiny, almost glabrous except for a few lateral short setae; weak alutaceous sculpture at base of notauli (Fig. 4A).Notauli complete with strong transverse carinae; median mesoscutal furrow short but distinct.Parapsidal signum short; parascutal sulcus present, internally carinate, being more distinct in anterior mesoscutum (Fig. 4B).Scutellar length 0.6 times that of mesoscutum in dorsal view.Scutellum alutaceous, shiny, completely areolate (Fig. 4A).Scutellar foveae rounded, smooth, without any internal carinae, basally defined by a carina; lateral pits of scutellar foveae present but superficial.Interfoveal line present.Circumscutellar carina complete, clearly defined, not dorsally projected at scutellum apex.Mesopleuron glabrous, shiny, with internally carinate transverse groove, presence of more or less extended coarse sculpture in anterior mesopleuron which may not be present in some cases.Mesopleural triangle densely covered by long hyaline hairs.Metanotal troughs densely pubescent.Propodeum heavily alutaceous, pubescent; central area longitudinally divided in two symmetrical areas by median carina; both areas are further divided by short transverse carinae in one big upper cell and two smaller cells.
Wings.Pubescent.radial cell of forewing closed, 2.7 times as long as wide.Marginal pubescence of forewing denser at apical third.
MetasoMa.Longer than head + mesosoma.Petiole about 2.0 times as long as metacoxa, smooth and shiny.Third metanotal tergum 2.1 times as long as fourth tergum in dorsal view.Fifth, sixth, seventh metanotal terga visible in dorsal view.Metanotal terga smooth and glabrous, punctate in anterior region of each tergum, more distinct from T4 to T7.

Taxonomic remarks
After examining large series of material, we have concluded that although most Anacharitinae species show a substantial degree of intraspecific variability in their body sculpture, the difference regarding the pronotal sculpture between Anacharis fergussoni sp.nov.and A. parapsidalis is greater than the variation among the individuals of each species, thus supporting A. fergussoni sp.nov.as a new species.

Biology
unknown.

Diagnosis
Species with incomplete notauli like A. immunis, from which can be distinguished by having a median mesoscutal furrow, a carinate sculpture in anterior mesoscutum and a strongly areolate scutellum (median mesoscutal furrow absent, mesoscutum completely smooth and scutellum smooth to tenuously areolate in A. immunis).This species is also morphologically similar to A. eucharoides, however it can be easily distinguished by the strongly areolate scutellum (scutellum alutaceous to rugose, never areolate, in A. eucharoides).

Etymology
The specific name makes reference to the distribution area of the type series.
CoLoration.Head, mesosoma and metasoma black.Mandibles yellowish brown with darker teeth.Antennae brown with black first segment.Legs yellowish brown, upper femur and coxae darker.Veins of wings brownish.
head.Triangular-shaped in anterior view.Face smooth, covered with abundant white setae.Width of head 1.2 times its height in anterior view and 1.9 times its length in dorsal view.Malar sulcus coriaceous, 0.6 times height of compound eye.Transfacial line as long as compound eye height.Diameter of toruli equal to torulus to compound eye distance but larger inter-toruli distance.Clypeus shortly defined, convex, margins densely covered by facial pubescence.occipital and postocular carinae absent.Compound eyes glabrous.In females, PoL:ooL:LoL ratio = 7:4.5:3,ocelli diameter being 2.5.Frons and occiput smooth covered by some scarce hyaline hairs, gena densely pubescent.
MesosoMa.Pronotum smooth, punctate, with shortly extended ridges from pronotal plate margin that become transverse carinae in ventral pronotum (Fig. 4E).Mesoscutal width 1.1 times its length in dorsal view.Mesoscutum smooth, shiny, almost glabrous except for a few short setae; weak carinate sculpture in region between notauli more distinct in anterior mesoscutum (Fig. 4D).Lateral region of mesoscutum smooth and glabrous, except for some peripheral short setae and punctures.Notauli weakly impressed, effaced in anterior mesoscutum, presence of weak internal carinate sculpture (Fig. 4D); median mesoscutal furrow short but distinct.Parapsidal signum tenuous, parascutal sulcus absent.Scutellar length 0.6 times that of mesoscutum in dorsal view.Scutellum alutaceous, shiny, completely covered by small areolate sculpture (Fig. 4D).Scutellar foveae triangular, smooth, without any internal carinae, basally defined by a carina; lateral pits of scutellar foveae elongated and deeply excavated.Interfoveal line present.Circumscutellar carina complete, clearly defined, not dorsally projected at scutellum apex.Mesopleuron glabrous, shiny, with internally carinate transverse groove alutaceous, presence of coarse sculpture in anterior mesopleuron.Mesopleural triangle densely covered by long hyaline hairs.Metanotal troughs coarse, densely pubescent.Propodeum coriaceous, pubescent; central area longitudinally divided in two symmetrical areas by a median carina; both areas are further divided by short transverse carinae.
Wings.Pubescent.radial cell of forewing closed, 3 times longer than wide.Marginal pubescence of the forewing denser at apical third.
MetasoMa.Longer than head + mesosoma.Petiole about as long as metacoxa (Fig. 4F), smooth and shiny.Third metanotal tergum 2.9 times as long as fourth tergum in dorsal view.Fifth, sixth and seventh metanotal terga visible in dorsal view.Metanotal terga smooth and glabrous, never punctate.

Biology
unknown.

Distribution
Palaearctic.only collected from Norway.

Taxonomic remarks
After examining the holotype, we found it has characters that fit within the genus Aegilips rather than Anacharis: petiole shorter than the metacoxa and coarsely sculptured, mesopleuron anteriorly carinated and without transverse mesopleural groove.For this reason we here recombine this species as Aegilips flavidicornis (Kieffer, 1910)  The known distribution of Anacharis immunis was divided between two areas before our study: one ranging from western Europe to the Caucasus while the other was limited to the russian Far East.The presence of A. immunis in Europe has now been consolidated with the addition of new citations from central Europe and the first specimens collected in southern Europe.The Asian distribution of the species is documented by two specimens only, both of them still limited to the Easternmost regions of Asia.This distribution pattern, divided into two areas, suggests a very plausible presence of the species across the whole Palaearctic region, but more specimens from Asia will need to be collected in the future to support this hypothesis.
regarding the distribution of the newly described species, Anacharis fergussoni sp.nov.has been found in southern and central Europe, while the distribution of A. norvegica sp.nov. is restricted to southern Norway.More interesting is the description of A. belizini sp.nov., the first species of Anacharis for the Indomalayan region, where the Anacharitinae record was previously limited to two species of Xyalaspis cited from Thailand (Mata-Casanova et al. 2014b).The new data on A. belizini sp.nov.makes the genus present in all regions and extends the knowledge of the anacharitines in the region.However, the Indomalayan anacharitines are still far from being well known.More research should be done in order to improve our knowledge of the Indomalayan anacharitines.
unlike Xyalaspis, which presents high interspecific variability in mesoscutal sculpture (Mata-Casanova et al. 2014a, 2014b), the variability in the sculpture of the mesoscutum in Anacharis is reduced to some weak carinae in some species.regarding their mesoscutal traits, species of Anacharis could be roughly divided between those species with complete and deeply excavated notauli usually internally carinate and those with incomplete notauli or so weakly excavated that they appear to be incomplete.In the case of Eurasian species, all species fall in the first category, except for A. immunis and A. norvegica sp.nov., which have tenuous notauli effaced in anterior mesoscutum (Figs 3C,4D).
More useful characters to distinguish between species of Anacharis are those related to the pronotum and the scutellum.When talking about the pronotum, most species of the genus have a smooth surface with only some short lower carinae; this is the case of A. antennata (Fig. 1B), A. eucharoides (Fig. 2D), A. immunis (Fig. 2E) -in which carinae are very reduced-and A. norvegica sp.nov.(Fig. 4E).Anacharis fergussoni sp.nov.represents the next step in pronotal sculpture, having a more extended lower carina, while the rest of the pronotum is slightly coarse (Fig. 4b).Anacharis parapsidalis (Fig. 1D) and A. belizini sp.nov.(Fig. 2B) represent the other side of the spectrum with a completely carinated pronotum.
regarding the scutellar sculpture, the studied species of Anacharis have a high degree of interspecific variability: Anacharis antennata and A. belizini sp.nov.have a smooth and shiny scutellum with no traces of carinae (Figs 1A and 2C,respectively); both species also present a short median scutellar carina at the apex of the scutellum, which can be interpreted as the remains of a more developed scutellar sculpture.The next degree in scutellar sculpture is seen in A. eucharoides, which has an alutaceous scutellar surface and shows traces of areolate sculpture near the circumscutellar carina (Fig. 2A-B).Anacharis immunis has a high degree of variability in scutellar sculpture, ranging from an almost smooth scutellum in some individuals (Fig. 2C) to having areolate sculpture in others (Fig. 2E-F), but their carinae are not as strong as in A. norvegica sp.nov., A. parapsidalis or A. fergussoni sp.nov. in which the surface is completely covered by strong areolated sculpture (Figs 4D,1C and 2C,respectively).The sculpture in the mesoscutum, however, is not constant: while A. parapsidalis and A. fergussoni sp.nov.have large cells across scutellar surface (Figs 1C, 2C), the scutellum in A. norvegica sp.nov. is divided into small cells (Fig. 4D).
The Palaearctic is the region where more hosts have been recorded for Anacharis, all of them belonging to subfamily Hemerobiidae (Neuroptera): Kierych (1984) listed Wesmaelius subnebulosus and Wesmaelius nervosus as hosts of Anacharis immunis; later, Fergusson (1986) listed Hemerobius micans, Wesmaelius betulinus and Wesmaelius subnebulosus as hosts of Anacharis eucharoides.There are 170 species of Hemerobiidae cited for the Palaearctic region (Makarkin 1995), so the species of Anacharis studied have plenty of potential hosts.More research should be done in order to unveil the life cycle of Eurasian Anacharis.
(Belizin 1961)aris flavidicornis to Aegilips, the synonymization of A. gracilipes with A. eucharoides and the description of A. belizini sp.nov., A. fergussoni sp.nov.and A. norvegica sp.nov., seven species of Anacharis are present in the Palaearctic and Indomalayan regions: Anacharis antennata, A. belizini sp.nov., A. eucharoides, A. fergussoni sp.nov., A. immunis, A. norvegica sp.nov.and A. parapsidalis.The number of valid species of Anacharis rises to twenty-two.For the Eastern Palaearctic species Anacharis antennata and A. parapsidalis are cited for the first time from Japan.Anacharis antennata maintains its known distribution area restricted to the northeastern Asian continent, from Japan to Khabarovskiy krai in russia.In the case of A. parapsidalis, previously known from Tajikistan in Central Asia only, the presence of specimens in Japan and one specimen from romania suggests a wide distribution area across the Palaearctic region.Anacharis eucharoides was mostly circumscribed to the Western portion of the Palaearctic region, with only one exemplar collected on the Asian continent(Belizin 1961); its known European distribution area previously comprised mostly Central and Eastern Europe.The new data presented in this work expands its known distribution to southern Europe and northwest Africa, with one specimen collected in Morocco.The current knowledge of the species distribution patterns suggests a Palaearctic distribution area centered in Central and Eastern Europe.