#NEXUS [!Export generated by Xper2 on 2025-03-17 11:42:04] Begin Data; Dimensions NTax=71 NChar=153; Format datatype=Standard Missing=? Gap=N; [Symbols="0 1 2 3 4 5 6 7 8 9";] [Options GapMode=Missing;] [CharStateLabels 1 DP1:_Dorsal_disc_scalation:/ few_thin_scales:/many_thin_scales:/thick_scales:/very_few_thin_small_scales_or_none, 2 DP3:_Dorsal_disc_scale_size:/ variable uniform, 3 DP4:_Dorsal_disc_scale_stereom_structure:/ no_tubercles:/with_smooth_tubercles, 4 DP5:_Central_primary_plate_relative_size:/ larger_than_disc_scales:/clearly_identifiable:/same_size,indistinguishable or absent, 5 DP6:_Primary_radial_plates_relative_size:/ larger_than_scales:/same_size, indistinguishable or absent, 6 DP7:_Primary_radial_plates_position:/ at_a_distance_from_CPP:/in_contact_with_CPP, 7 DGS1:_Dorsal_disc_covering_nature:/ without_granules_spines:/with_granules_only:/with_both_granules_and_spines:/with_spines_only, 8 DGS2:_Dorsal_disc_granules or spines_extension:/ sparse_all_over_with_underlying_plates_or_scales_visible:/restricted_to_margin:/forming_dense_cover_completely_hiding_underlying_plates_scales_possible_exception_radial_shields, 9 DGS5:_Radial_shield_granulespine_covering:/ naked:/at_least_partly_covered, 10 DGS6:_Dorsal_disc_granules_spines_differentiation:/ uniform:/modified_e.g._enlarged_at_disc_edge, 11 DRS3:_Radial_shields_in_articulated_disc_plating_length:/ less_than_one_third_of_the_disc_radius:/between_one_third_and_half_of_the_disc_radius:/more_than_half_of_the_disc_radius, 12 DRS4:_Radial_shield_pairs_in_articulated_disc_plating_proximity:/ completely_separated:/separated_distally ;separated_proximally:/in_contact_over_entire_length, 13 DRS5:_Radial_shield_shape:/ scalene_triangular:/isosceles_triangular_to_pear-shaped:/half-circle:/bar-like, 14 DRS6:_Radial_shield_abradial_edge_shape:/ entire:/with_extensions_regular_outline_enlarged:/incised;irregular, 15 DRS7:_Radial_shield_exposure:/ central_part_of_RS_to_almost_entire_RS_exposed:/distal_portion_of_RS_exposed:/distal-adradial_portion_of_RS_exposed:/completely_covered_by_scales, 16 DRS9:_Muscle_attachment_area_on_RS_shape:/ round_flat:/crater-like_depression_with_proximal_protruding_edge-rim:/large_depression_(Gorgonocephalus), 17 DRS10:_RS_articulation_shape:/ one_condyle:/two_condyles-spurs_with_depression_between_them:/no_special_structure:/smooth_patch, 18 DRS11:_RS_distal_edge_shape:/ groove_and_lip-shaped_edge:/other_shape, 19 DRS12:_Conspicuous_pore_in_inner_side_of_RS:/absent:/present, 20 VI2:_Ventral_interradii_covering:/ with_granules/with_spines/naked with_tubercles, 21 VI3:_Genital_slit_length:/ shorter_than_half_interradius; longer_than_half_interradius_or_divided_into_two_openings; lost, 22 ORGP1:_Oral_GP:/absent; present, 23 ORGP2:_Oral_GP_shape:/bar-like; oval; L-shaped; rectangular; comma-shaped; drop-shaped, 24 ORGP3:_Rectangular_oral_GP_details:/with_folded_edge; convex; with_processes, 25 ORGP4:_Oval_oral_GP_details:/pyramid-like_raised; with_distal_notch; flat, 26 ADGP1:_Adradial_GP_shape:/bar-like; multi-layered; stout_half_as_wide_as_long; blade-like_straight_excluding_distal_tip_curving_dorsal; wrench-shaped; comma-shaped; cleaver-shaped; cup-shaped, 27 ADGP2:_AdGP_to_RS_articular_structures_shape:/single_condyle; 2_condyles; flat; condylar_process, 28 ADGP3:_Barlike_adGP_subtypes_shape:/distally_inflated; distally_not_inflated; club-like, 29 ADGP4:_Additional_structures_on_adGP_shape:/patch_of_different_stereom; process, 30 ABGP1:_Abradial_genital_plate_relative_length:/shorter_than_half_the_adradial_plate_length; longer_than_half_the_adradial_plate_length; as_long_as_adradial_plate, 31 ABGP2:_Abradial_genital_plate_shape:/strong_curved_scale; thin_as_wide_as_long; sabre_or_sickle-shaped; twisted; triangular_to_pear-shaped; blade-like_straight; trapezoid_to_rectangular; scoop-shaped; cup-shaped; thin_curved_scale, 32 ABGP3:_Shape_of_adradiodistal_articular_structure_of_abradial_genital_plate:/straight_or_convex; concave; fused_to_adGP, 33 ABGP6:_Abradial_genital_plate_covering:/fully_covered; exposed, 34 ABGP7:_AbGP_plate_papillaegranules_nature:/disc_granules; papillae, 35 ABGP8:_AbGP_papillae_extension:/restricted_to_ventral_disc; extending_to_laterodorsal_disc_edge, 36 ABGP9:_Ventral_genital_papillae_shape:/granule-like; spine-like; block-like, 37 ABGP10:_Conspicuous_perforation_in_adradial_surface_of_abGP:/absent; present, 38 ABGP11:_Longitudinal_groove_in_abradial_surface:/absent; present, 39 ABGP12:_distalmost_genital_papillae_on_dorsal_disc_shape:/block-like; spine-like, 40 ABGP13:_Abradial_GP_crosses_adradial_GP:/no; yes, 41 MOAS1:_Oral_shield_length:/covering_less_than_one_third_of_interradius; longer_than_one_third_of_length_of_interradius, 42 MOAS2:_Oral_shield_shape:/longer_than_wide; as_long_as_wide; wider_than_long, 43 MOAS3:_Madreporite_size:/similar_to_other_oral_shields; larger_than_remaining_oral_shields, 44 MOAS4:_Madreporite_inner_side_structure:/1–2_openings; meshwork_of_openings; several_pores, 45 MOAS5:_Madreporite_inner_opening_position:/proximal; central; distal; centrodistal, 46 MOAS6:_Adoral_shields_proximity:/separated; meeting_proximal_to_oral_shield, 47 MOAS8:_Oral_shield_proximal_portion_shape:/evenly_convex; obtuse_angle_with_straight_to_convex_sides; obtuse_angle_with_concave_sides; acute_angle_with_straight_to_convex_sides; acute_to_right_angle_with_convex_to_straight_sides, 48 MOAS9:_Oral_shield_distal_portion_shape:/evenly_convex; square; with_narrower_distalward_projection, 49 MSP1:_Second_tentacle_pore_position:/deep_inside_mouth_slit; shallow_and_oblique, 50 MPaT1:_Lateral_papillae_arrangement:/single_row_along_jaw_edge; multiple_rows_covering_jaws, 51 MPaT2:_Ventralmost_tooth_position:/exposed; obscured_by_tooth_papillae_or_infradentals, 52 MPaT3:_Buccal_scale_shape:/smaller_papilla_Ophiura; pointed_wide_higher_on_oral_plate_Amphiura; single_wide_papilla_Amphilepis; lost, 53 MPaT4:_infradental_papillae_position:/lateral_close_to_dental_plate; lateral_at_distance_from_dental_plate; below_teeth, 54 MPaT5:_Adoral_shield_spine_shape:/spiniform; scale-like_or_like_other_papillae; scale-like_much_largerwider_than_other_papillae, 55 MPaT6:_Second_degree_ASS_position:/at_oral_plate; at_2nd_tenacle_pore, 56 MPaT7:_Additional_lateral_papillae_shape:/block-shaped; rounded; paddle-shaped; spiniform, 57 MPaT8:_Teeth_shape:/spine-like_thin_and_sharply_pointed; with_round_or_slightly_pointed_tip_but_never_spinelike; with_square_tip, 58 MPaT9:_Infradental_papillae_relative_size:/similar_to_other_oral_papillae; larger_than_other_oral_papillae, 59 MPaT10:_Second_degree_adoral_shield_spine_number:/single; multiple, 60 MPaT11:_Lymans_ossicle_position:/oblique_in_mouth_slit; in_line_with_other_papillae, 61 MPaT12:_Tooth_papillae_number:/2; >2, 62 MPaT13:_Accessory_oral_papillae:/absent; present, 63 MDP1:_Dental_plate:/entire; several_pieces, 64 MDP2:_Dental_plate_geometry:/equal_width_all_over; ventral_half_widest dorsal_half_widest, 65 MDP3:_Tooth_sockets_pattern:/single_column_throughout; multiple_columns_or_cluster_on_max_half_of_plate; multiple_columns_throughout, 66 MDP5._Tooth_sockets_predominant_shape:/surrounded_by_a_more_or_less_continuous_protruding_ring; simple_opening surrounded_by_separate_weakly_protruding_knobs_andor_ridges; surrounded_by_strongly_protruding_knobs_andor_ridges, 67 MDP7:_Tooth_sockets_depth:/depression_or_perforating_DP_without_septum; at_least_some_perforating_DP_with_septum, 68 MOP1:_Oral_plate_shape:/as_high_as_long_or_higher; longer_than_high, 69 MOP2:_Abradial_muscle_fossa_shape:/large_well_defined_flange; central_depression, 70 MOP3:_Abradial_muscle_attachment_area_stereom_structure:/normal_stereom; rib-like_branching_structures, 71 MOP4:_Adradial_muscle_attachment_area_position:/ventral_lining_ventral_or_ventrodistal_edge_of_articulation_area; in_middle_position_vertical_and_lining_less_than_two_thirds_of_distal_edge_of_adradial_articulation_area; in_middle_position_vertical_and_lining_more_than_two_thirds_of_adradial_articulation_area; with_large_dorsal_spoon-shaped_depression; flange_over_most_of_lateral_surface_with_striations, 72 AG2:_Integument_of_arms_structure:/naked; bearing_granules, 73 AG3:_Arms_shape:/simple; branching, 74 AVP3:_Proximal_VAPs_in_articulated_plating_position:/separated_by_lateral_arm_plates; potentially_in_contact, 75 AVP4:_Distal_edge_of_proximal_VAPs_in_macerated_plate_shape:/convex_to_straight; concave_or_incised, 76 AVP5:_Proximal_edge_of_proximal_VAPs_macerated_shape:/convex_to_straight; concave_or_incised, 77 AVP6:_Distal_portion_of_proximal_VAPs_macerated_dimensions:/as_wide_as_proximal_portion_or_narrowerM; wider_than_proximal_portion, 78 AVP7:_Lateral_edge_of_proximalmost_VAPs_macerated_shape:/not_incised; with_clear_incisisons_or_notches_for_tentacle, 79 AVP8:_Edge_of_tentacle_notches_with_sockets-articulations_for_tentacle_scales_e.g._Ophiomyces:/absent; present, 80 AVP10:_Proximal_VAPs_ornamentation:/without_conspicuous_ornamentation; with_tubercles_or_striation, 81 AVP12:_Spurs_at_proximal_edge_of_VAPs_macerated:/absent; present, 82 ADP2:_DAPs_per_segment_number:/single; multiple; none, 83 ADP3:_Proximal_DAP_series_position:/separated; in_contact, 84 ADP4:_Proximal_DAPs_shape:/fan-shaped; trapezoid_with_smooth_proximal_edge; oval_semicircular, 85 ADP8:_Proximal_DAPs_ornamentation:/without_conspicuous_ornamentation; tuberculous; with_striation, 86 ADP10:_Spurs_at_proximal_edge_of_DAP_macerated:/absent; present, 87 ADP11:_Accessory_dorsal_arm_plates:/absent; present, 88 AS1:_Arm_spine_position:/lateral; at_prox._segments_only_on_ventral_side_of_arm, 89 AS2:_Arm_spines_orientation:/predominantly_parallel_to_arm_axis_adpressed; predominantly_erect_standing_perpendicular_to_arm, 90 AS3:_Longest_spine_length:/shorter_than_half_a_segment; between_half_a_segment_and_one_segment; 1-2_segments; longer_than_2_segments, 91 AS4:_Arm_spine_interior_structure:/massive; with_lumen, 92 AS5:_Arm_spine_surface_ornamention:/smooth; with_lateral_thorns; with_scalelike_tubercles, 93 AS7:_Arm_spine_cross_section_shape:/round; laterally_flattened, 94 AS8:_Tip_of_arm_spines_shape:/blunt; pointed, 95 AS10:_Hookshaped_arm_spines_position:/at_no_segments; only_at_distal_segments; at_proximal_and_distal_segments, 96 AS11:_Hookshaped_spines_shape:/regular_spines_with_bent_tip_and-or_saw-toothed_edge; true_hyaline_hook; both_modified_regular_spines_and_true_hooks, 97 AS13:_Arm_spines_size_pattern:/ventralmost_spines_longest; median_spines_longest; dorsalmost_spines_longest; all_equal, 98 AS14:_Arm_spines_at_proximal_to_median_arm_segments_number_changes:/decreasing_distalwards; constant, 99 ATS3:_Tentacle_scales_shape:/operculiformround_to_oval leaflike_blunt spinelike ventral_spine_closing_tentacle_opening flat_elongated, 100 ATS4:_Tentacle_scales_size:/not_accurately_closing_tentacle_pore accurately_closing_tentacle_pore_or_larger, 101 ATS5:_Tentacle_scales_ornamentation:/without_longitudinal_striation with_longitudinal_striation, 102 ATS6:_Tentacle_scale_placement:/only_at_LAP at_both_LAP_and_VAP only_at_VAP, 103 AV1:_Dorsodistal_muscular_fossae_distalwards_projecion:/not_projecting far_from_distal_edge_of_zygocondyles_e.g._Ophiacantha almost_beyond_zygocondyles_e.g.._Ophiodoris beyond_zygocondyles_e.g._Ophiothrix, 104 AV2:_Lateral_saddle_between_muscular_fossae_structure:/with_single_ridge with_multiple_knobs_e.g._Gorgonocephalus, 105 AV4:_Zygocondyles_in_proximal_vertebrae_position:/nearly_parallel dorsalwards_converging, 106 AV6:_Zygosphene_fused_with_pair_of_zygocondyles_size:/not_projecting_beyond_ventral_edge_of_zygoc._or_projecting_beyond_ventral_edge_of_zygoc._with_projecting_part_shorter_than_zygoc.; projecting_beyond_ventral_edge_of_zygoc._with_projecting_part_as_long_as_zygoc.; projecting_beyond_ventral_edge_of_zygoc._with_projecting_part_longer_than_zygoc., 107 AV7:_Needlelike_thorns_in_dorsal_groove:/absent; present, 108 LAPG1:_LAPs_position:/only_lateral; arched, 109 LAPG3:_LAPs_with_constriction:/absent; present, 110 LAPG4:_Ventroproximalwards_projection_of_ventral_portion_of_LAP:/absent; present, 111 LAPG5:_Ventralwards_projection_on_ventrodistal_tip_of_LAP_e.g._Ophiopallas:/absent; present, 112 LAPG6:_Dorsal_edge_distalwards_ascending:/absent; present, 113 LAPG7:_First_LAPs_shape:/like_following_LAPs; winglike_folded; elongated_blade-like, 114 LAPO2:_Outer_surface_trabecular_intersections_shape:/not_protruding; protruding_to_form_only_knobs_approx._the_same_size_as_stereom_pores; protruding_to_form_knobs_larger_than_stereom_pores_on_most_of_outer_surface_of_LAP; protruding_to_form_knobs_larger_than_stereom_pores_on_small_part_of_outer_surface_of_LAP, 115 LAPO8:_Outer_surface_structure:/normal_stereom; vertical_striation_formed_by_merged_knobs; vertical_striation_formed_by_regular_ridges; with_pointed_thorns, 116 LAPO10:_Distalwards_pointing_scale-like_structures_on_outer_surface_stereom_presence:/absent; present, 117 LAPPE1:_Proximal_edge_of_outer_LAP_lined_by_discernible_band_of_different_e.g._more_finely_meshed_stereom_structure_position:/no_discernible_band_of_different_stereom; only_in_central_part; over_most_of_the_proximal_edge, 118 LAPPE2:_Spurs_on_prox._edge_of_outer_LAP_surface_other_than_ventroproximal_one_size:/more_than_two_small_shorter_than_14_of_LAP_width; 1_or_2_large_wider_than_14_of_LAP_width; 1_or_2_small_shorter_than_14_of_LAP_width, 119 LAPPE3:_Oblique_elongated_spur_on_ventroproximal_tip_of_outer_surface_of_LAP_presence:/absent; present, 120 LAPPE4:_Proximal_outer_surface_edge_of_LAP_extension_central_part_extension:/not_protruding; protruding, 121 LAPPE8:_Proximal_edge_of_outer_LAP_surface_with_horizontal_striation_extension:/restricted_to_small_area_e.g._between_spurs; along_most_of_the_edge, 122 LAPSA1:_Spine_articulations_position:/on_same_level_as_remaining_outer_surface; on_elevated_portion_bordered_prox._by_ridge; on_elevation_not_bordered_by_ridge; in_notches_of_distal_LAP_edge, 123 LAPSA4:_Spine_articulation_separation_from_distal_edge:/separated_from_distal_edge_by_the_usual_outer_surface_stereom; separated_from_distal_edge_by_a_thin_projection_of_the_distal_LAP_portion_e.g._Ophiomyces; directly_adjacent_to_the_distal_edge_of_the_LAP, 124 LAPSA5:_Spine_art._series_extension:/restricted_to_ventral_or_central_portion_of_distal_LAP_edge; arranged_over_entire_distal_LAP_edge, 125 LAPSA6:_Spine_art._size:/dorsalwards_increasing; ventralwards_increasing; middle_spine_art._larger; all_similar, 126 LAPSA7:_Spine_articulations_distance:/dorsalwards_increasing; ventralwards_increasing; equidistant; only_2, 127 LAPSA8:_Nerve_and_muscle_openings_separation:/by_small_ridge_if_at_all; by_large_prominent_ridge_or_regular_stereom, 128 LAPSA9:_Nerve_opening_size:/smaller_than_muscle_opening; approx._as_large_as_muscle_opening, 129 LAPSA10:_Articular_structures_shape:/like_surrounding_stereom; elevated_lobes, 130 LAPSA11:_When_dorsal_and_ventral_lobes_absent_prox._edge_of_muscle_opening_structure:/even; denticulate, 131 LAPSA12:_Dorsal_and_ventral_lobes_connection:/simply_separated separated_by_one_or_several_knobs_or_by_denticulate_stereom; merged_at_their_proximal_tips_by_smooth_connection, 132 LAPSA13:_Dorsal_and_ventral_lobe_size:/one_lobe_clearly_larger_than_the_other; equal-sized, 133 LAPSA14:_Dorsal_and_ventral_lobes_orientation_to_each_other:/shifted_e.g._Ophiacantha; parallel_e.g._Amphiura, 134 LAPSA15:_Dorsal_and_ventral_lobes_shape:/at_least_one_lobe_bent; straight, 135 LAPSA16:_Dorsal_and_ventral_lobes_stereom_structure:/with_perforations; massive, 136 LAPSA17:_Lobes_orientation_on_plate:/nearly_horizontal; tilted; nearly_vertical, 137 LAPSA19:_Sigmoidal_fold_expression:/weakly_developed; fully_developed, 138 LAPSA20:_When_dorsal_and_ventral_lobes_absent_muscle_opening_encompassed_by:/simple_stereom_e.g._Euryale; poorly_defined_circular_elevation_e.g._Asteronyx; and,or_vertical_ridge_distally_and_wavy_ridge_prox._e.g._Gorgonocephalus; vertical_mouth-shaped_sharply_defined_elevation, 139 LAPSA21:_When_dorsal_and_ventral_lobes_absent_orientation_of_ridge_bordering_muscle_opening:/vertical; oblique, 140 LAPSA22:_when_dorsal_and_ventral_lobes_absent_shape_of_ridge_distally_bordering_muscle_opening:/slender; thick_lip-shaped_and_strongly_protruding, 141 LAPTP1:_Tentacle_opening_beyond_first_segments_under_disc_shape:/notch; within-plate_perforation, 142 LAPTP2:_Tentacle_notch_direction:/ventralwards; ventralwards_to_ventrodistalwards; distalwards_close_to_horizontal_midline_of_LAP, 143 LAPTP3:_Tentacle_notch_externally_lined_by_narrow_groove:/no; yes, 144 LAPTP4:_Inner_side_of_notch_with_horizontally_stretched_stereom:/no; yes, 145 LAPI1:_Inner_side_of_LAP_main_structure:/more_or_less_continuous_ridge; two_separate_rarely_merged_central_knobs, 146 LAPI3:_Ridge_on_inner_side_of_LAP_shape:/entire; with_ventral_tip_of_ventroproximalwards_pointing_part_of_ridge_separated_from_remaining_ridge; with_separate_knob_on_ventral_tip_of_LAP; ridge_separated_into_two_halves, 147 LAPI5:_Ridge_on_inner_side_og_LAP_stereom_structure:/same_stereom_as_remaining_inner_surface_of_LAP; more_compact_or_more_densely_meshed_stereom, 148 LAPI6:_Ridge_on_inner_side_of_LAP_shape:/without_major_kink_and_with_tongue-shaped_dorsal_tip; with_kink_between_dorsoproximalwards_pointing_dorsal_portion_and_ventroproximalwards_pointing_ventral_portion; with_two_kinks_and_dorsal_tip_with_ventroproximalwards_pointing_projection; with_two_kinks_and_dorsal_kink_with_ventroproximalwards_pointing_projection, 149 LAPI9:_Two_central_knobs_on_inner_side_of_LAP_shape:/simple; with_a_knob; with_a_ridge, 150 LAPI10:_Additional_dorsal_structure_on_inner_side_of_LAP_merged_with_prox._one_of_two_central_knobs:/no; yes, 151 LAPI7:_Large_dorsal_contact_surface_with_opposite_LAP:/absent; present, 152 LAPI8:_Large_ventral_contact_surface_with_opposite_LAP:/absent; present, 153 LAPI15:_Perforations_on_inner_side_of_LAP_shape:/small_or_inconspicuous; vertical_row_without_furrow; vertical_row_with_furrow; single_large_and_conspicuous ;] Matrix Aganaster_gregarius 0100000N0N00100????21????6(01)N????????00??010??13200????????????000??11000000011000000000000?000??10000?001101011001002?10?000321101NNNNNNN11010100010NN000 Acrocnida_brachiata 1000000N0N0001000003011N200002210???01??011??1120001210?21???0020310013001110100001100001110100N004001000101010000001?01?02110001N011110?NNN01001NNN11003 Actinozonella_texturata 2010000N0N100000111200???22N?2001???00?0010??1300002010?1001?00202011020000011010000101000?0010N300001000001000002002210?02032001N011110?NNN01000100NN000 Amphilepis_norvegica 1100000N0N20002000021????00101200???00?002101132000?0???10???0000201100000101100000200001200010N10????101101010001001?00?01110001N111110?NNN01001NNN0N000 Amphilimna_olivacea 11000?300011002000011132N00100310???00?0011021401000200?100??0020201102001001100101200001200110N002012201201011012001?00?02120011N111110?NNN00001NNN0N001 Amphiophiura_insolita 0000000N0N021000301210???22N?26011100010110001201010111?101?000110011000010000N10011100000?0010N110001200201000001002?10?021100100NNNNNNN11100000301NN000 Amphiophiura_latro 2101010N0N032000301210???22N?26011110010100001311010111?101?000110011010000000N0001100000000110N210001201101000002002?10?021320?01NNNNNNN11102000000NN000 Amphipholis_squamata 11001N0N0N2320200002111N?001?2500???00?0011??142000?220?210??00200100030000000N0000000001010010N300101000101010001001?00?02132001N011110?NNN01001NNN0N003 Amphiura_atlantidea 1100000N0N10012????211???000022?0???00?0021??112000121??21???00202100130011001000010000012?00120100000000101010001002?00?02112011N011110?NNN010???????00? Amphiura_chiajei 1100000N0N1021200002114NN00012210???00?101103112000121??21???00203100130011000N0001000001210010N100001201201010001001?00?02110011N011110?NNN01001NNN11003 Amphiura_filiformis 11001N0N0N12112????2110NN00012210???00??011??102000120??21???00203100030010000N0001000001211012110????000001010001001?00?02110001N011110?NNN01001NNN0N000 Aplocoma_agassizi 11001N100010101?100011???000?2600??????0000??132000?????0?????000001100000001100001000000100000N000100000001011001002200?00112001N2000010NNN01000010NN001 Aspiduriella_scutellata 000001100023100????00???????????????????010??1000000?1??10?????????11000000010N00000000000?0100N300001000001010000000?10?021301100NNNNNNN01101100001NN000 Aspiduriella_streichani 000001100022100????00??????????00??????0001??1000110???110?????????????0010010N00010000000?0100N3000?1000001010000000?00?001301100NNNNNNN011010?0001NN00? Asteronyx_loveni 3??11N0N0N201000210200???33N?1421???00?0011111301000???30???01002000102001?????002?NNN0110011022203000000?00000000000?00?021311000NNNNNNN0??01000300NN001 Breviturma_brevipes 11011N1110001210001011???001?2210???11?0000??00000100101200110001310004001001101001210001202100N000100100101010002002?00?20130001N2000011NNN01000213NN003 Breviturma_krohi 01011N1110001210001011???001?2200???11?0000??00000100101200110001310004001001101001210001200100N000100100101010002002?00?20130001N2000011NNN01000213NN003 Eirenura_papillata 1101??2111121020000011?????????00???????100??1411000111?101???000101100001101101000100000001100N001011101001011002112?10101112001N200012?NNN01000011NN001 Euryale_aspera 3??11N1010201002311200???22N01401???00?001011130110????11????11000001021110100N002?NNN0110?10021013000010?00000000000?00?020331100NNNNNNN0??11000300NN001 Gorgonocephalus_caputmedusae 3??11N301020100231?100???020?1500???00?0021111111110???30???11102000102111?????00112000110011022003000010?00010000000?00?020311000NNNNNNN00001000300NN001 Hemieuryale_pustulata 2010000N0N2010003002011N?52N0???????00?0121??110000?21??10?1?0020201101001001101011?001101?0000N000100000?00010002001210002113001N111100?NNN01000100NN000 Histampica_duplicata 1000000N0N1020200102111N?43N11510???00?001101140000001011001?0000201101001001100000000001210000N200100101001010002001?01002120011N001010?NNN01001NNN10001 Inexpectacantha_acrobatica 010001101000101????011??????????0??????0111??1320000010?1000000002011000000011010000100013?10111201110100?01100002002200120100001N2000011NNN01000112NN002 Melusinaster_allissawhitegluzae ???????????????????????????????????????0????????????????????????????????0????????????????????????0????000?01010000000?00?001301100NNNNNNN00001010001NN000 Ophiacantha_abyssicola 11011N311000301?210111???00110200???11?002101122000?0103100??000020010100010???0000000001311010N200100100001100003202?00?10130001N2000011NNN01000112NN002 Ophiacantha_bidentata 11011N311000301021111130N00110200???11?002002120000?0103100??0000201100000001101000000001311010N200110100001100003202?00?10100001N2000011NNN01000112NN002 Ophiactis_abyssicola 10011N30002000000002115NN40N11010???00?0021011200001?2012?0??0010200002000001100000000001211000N200100100101010002001?01002112001N001010?NNN01001NNN10003 Ophiactis_profundi 11001N0N0N200000000211???40N11010???00?0011??1200001?2??1????00002000020010001000000000012110010210100100101010001001?01?02112001N001010?NNN01001NNN10003 Ophiarachna_incrassata 11011N111000121????0111N?01001500???11?0020??102001?0201100100101001100001001101101111001302010N0011?0000001011002002110?32112001N2000010NNN01000210NN001 Ophienigma_spinilimbatum 1000003100002020000211???40N02011???00??020030200000010?200??0020201101001101101001010001200010N201110100001010002001?01?02120001N001010?NNN01011NNN10001 Ophiocamax_vitrea 10011N30002312000011114NN00002510???10?002001112111?0013001010001000100000011101000010001311010N201100100?11100003302200110120001N2000011NNN00010011NN000 Ophiochiton_fastigatus 1001000N0N10002????211???63N02500???00?0021??142000?020?1000?00202011010010011000012000012?0010N200101201101011001001?10102120001N111100?NNN01000113NN003 Ophiocopa_spatula 11001N10000310003002111N?00011910???10?011001132000?0201200?00100201100000001101000010001311100N200110100001110003202200?10132001N2000011NNN01010112NN002 Ophiocoma_echinata 11011N111000121????011???001?1200???11?0010??100001?020?200110001310004001001101001111001302010N200100000001010002002?10?22100001N2000011NNN01010213NN003 Ophiocoma_scolopendrina 11011N11100012200010112NN00111200???11?000002000001?01??20?110001310004001001101001010001302000N200100000101010002002?00?20100001N2000101NNN01000213NN003 Ophiocomina_nigra 11011N11100010101012115NN01111910???10?002102120001?0203200010011001100001101100001000001301110N201110100001110001202?00?10100001N2000011NNN01010113NN002 Ophiocten_sericeum 1000000N0N00100????210???001018011110010100??14010001110101?0?0020111000000010N0001100001200010N211000100201000001202?10?310311000NNNNNNN10002000310NN001 Ophioderma_longicaudum 11011N11100012101010114NN01010600???11?002002102000?0101200101101001100001001101111111000002100N000100001001011002002110032112001N2000010NNN01000210NN001 Ophioderma_africanum 11011N1110001110101011???01010600???11?001002102000?0101200100111001100001001101101111000002100N000100001001011002002110032112001N2000010NNN01000010NN001 Ophiodoris_malignus 11001N0N0N000020111211???60N0251000000?0010??0020000020?2001?0000201101001001100001200001200010N100111201101010002001?10002120001N101100?NNN01010113NN003 Ophiolepis_superba 2000000N0N1010001112111N?60N02411???10?0000??1320000220?2001?0020201101001001100011100100000010N000100001001010001002110032120011N111100?NNN01000200NN000 Ophiolepis_variegata 2000000N0N1010001102111N060N02401???10?0000011320000220?2001?002000110100100110011110110000001102001?0001001010001002110032022001N111100?NNN01000000NN000 Ophioleuce_seminudum 100100111000002????011??????????000000??000??132001?010?000?10010101100000001101001121000101010N300110101001011002112200101113001N200012?NNN01010111NN000 Ophiernus_vallincola 000???100010100?110211???00111510???00?0010010121000010?100?00000101101001001100001111000001010N300000200001100000002200101032001N210011?NNN00010000NN000 Ophiolimna_antarctica 11011N1110103010200011???02111210???00?00200211000100203100?00011201100001001101001020001310010N201110000001100001202?00?10102001N2000111NNN01000112NN002 Ophiolimna_bairdi 11011N2110003010200011???02111200???00?01100212000000203100?00010201100000001101000020001211010N201110100001100001202?00?10100001N2000111NNN01000112NN002 Ophiolycus_purpureus 11011N30100030302101111N202110500???00?002102122101????31???0?011001100001101100?0110?0012010011102010101001111000002?10000100001N2000010NNN01010111NN001 Ophiomusa_lymani 2000000N0N201000000200???03012601???00?0100011310000120?100??1020001100000001100000000000000000N000100001101000101002010?001321101NNNNNNN11110100010NN000 Ophiomyces_delata 01011N30?0?????????110???3?N12500???00?000021032010?00031?0?1?010101101000101111000000001201110N100011100001011001202?10?01112001N200012?NNN01000111NN002 Ophiomyxa_pentagona 31011N0N0N00311?3112114NN02111500???10?001102100000?01?020?0??100001100001101100001100000001001010????000001011000000210?00112011N2000010NNN01000210NN001 Ophiomyxa_serpentaria 31011N0N0N00321?3112114NN02110500???10?002102010000?01?021?0??1000011000010001000011000002010110004000000001011000000210?00112011N2000010NNN01000210NN001 Ophionereis_porrecta 10011N0N0N000020000211???63N025100000011010020020000220?2001?0020311003001001101011210101200010N100101300101010002001?10?02120001N101100?NNN01010113NN003 Ophionereis_reticulata 10011N0N0N0001200002111N263N0251000000?1010020020000220?2001?002001000300110010000110010120010??100100300101010002001?10?02112001N101100?NNN0101000?NN003 Ophiopallas_paradoxa 10011N11100010200000111N030N12010011001010001030000?02??00??100200011000010011010011210001010111100100101001011002112000101120001N200012?NNN01010111NN000 Ophiopholis_aculeata 10101N21111020200001111N230N00010???00?0021011100010?20?2?0?100113100130011101010012100012010111100010301001010002001?01?22120011N201001?NNN01001NNN21003 Ophioplax_lamellosa 10001N0N0N0020200000111N?60N0251000000?0110021320000220?1001??000201101001001100001200001200000N100101201201010001001?10002120001N101100?NNN01010113NN003 Ophiopleura_borealis 10001N0N0N003010000200???0(03)00270010000?0000011400000111?001?010000011000000010N1001110000000010N110000101201000001002?10?320101101NNNNNNN10102000300NN000 Ophiopsila_guineensis 11011N0N0N1030200002110NN00012210???00?0011021020010?1??2???10020310003001101100001200001200100N004110201101011000001?00?02110011N111110?NNN00011NNN0N001 Ophiopus_arcticus 1000000N0N001010000220???40N0???????00??020031300000020?200??0000200102000001101000010001200010N301100100001010001001?00002130001N011010?NNN01011NNN0N003 Ophiopyrgus_saccharatus 2010010N0N121000311210???22N124111110010100??1301000111?101?0?000001101000001101000010000001010N310011200101000002102?001021331101NNNNNNN01001000000NN000 Ophiopyrgus_wyvillethomsoni 2100010N0N01120?010210???22N?241010200?0100??1101000121?100?0?0100011010000011000000000010?0000N310000000101000000002?00?021330100NNNNNNN0??1???0011NN000 Ophioscolex_glacialis 31011N0N?N?????????2111N10?110500???10?002102100101??0??0???1?01110110000100110002?NNN001201010N20????001001111000002?10102100001N201000?NNN00010111NN000 Ophiotholia_spathifer 1101??0N?N?????????210???3?N?2500???00?0000??032010?00031?0010010101101000001110000000001201010N100011100001111001202?10001112001N201010?NNN00000111NN001 Ophiothrix_fragilis 11100130012000200001111N100011010???00?0020010100?13????1???100113100130011111010011100013010121102000301001010002001?01?22120011N201001?NNN01001NNN20003 Ophiotreta_valenciennesi 11011N1110101011001011???00010910???10?0010??132001?01012000101110011000010011000010000013?1110N200110101001110001202?00?10100001N2000011NNN01000113NN002 Ophiozonella_longispina 1010000N0N101000000211???53N02711???00?0010011300000020?2000?0020211102001001101001111101200010N200100001001011002002210032133011N111100?NNN01000200NN000 Ophiura_ophiura 1000000N0N101000100210???010018111120010100011401010111?101?110020011000000010N0001110000100100N210001101201000022102?10?320011101NNNNNNN10002000300NN001 Ophiura_sarsii 1000000N0N101100100210???010018111120010100011401000111?101?110120011000000010N0001110000200100N210001101201000022102?10?320021101NNNNNNN10002000300NN000 Palaeocoma_milleri 100???110020100????010???00000811112??00100??1310010?20?10010?001101100001101100001100000000100N2000010000010100010020100321311100NNNNNNN10001100010NN111 Sigsbeia_murrhina 2010000N0N201?0????201???22N????????00?0020??1100002010?100??0?????????001011101011010110000100N3101?0001?01010002001110002033001N111100?NNN01000100NN000 Warenophis_andersi 00111N0N0N23000????211??????????0???????021??1101013110?210?00?????????000001100001000001201100N200000000101110001002200?20102001N2000020NNN01010210NN001 ; outgroup Aganaster_gregarius; lset rates=gamma; lset coding=variable nbetacat=5; prset symdirihyperpr=fixed(infinity) ratepr=variable; prset brlenspr=unconstrained:exponential(1.0) shapepr=exponential(1.0); mcmc ngen=9000000 samplefreq=1000 printfreq=10000 starttree=random; End; End;