Calcetas, O.A. 2023. Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896. European Journal of Taxonomy. 890, 1 (Aug. 2023), 184-203. DOI:https://doi.org/10.5852/ejt.2023.890.2261.