Rossini, M., Vaz-de-Mello, F. Z., & Zunino, M. (2018). Toward a comprehensive taxonomic revision of the “hirculus” group of American Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae). European Journal of Taxonomy, (432). https://doi.org/10.5852/ejt.2018.432