Calcetas, Orlando A. 2023. “Three New Species of the Genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and Designation of a Neotype for L. Semperi Brenske, 1896”. European Journal of Taxonomy 890 (1):184-203. https://doi.org/10.5852/ejt.2023.890.2261.