Calcetas, O. A. (2023) “Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896”, European Journal of Taxonomy, 890(1), pp. 184-203. doi: 10.5852/ejt.2023.890.2261.