European Journal of Taxonomy <p>The <em>European Journal of Taxonomy</em> is a peer-reviewed international journal in descriptive taxonomy, covering the eukaryotic world. Its content is fully electronic and <a href="">Open Access</a>. It is published and funded by a <a href="">consortium</a> of European natural history institutions. Neither authors nor readers have to pay fees. All articles published in <em>EJT</em> are compliant with the different nomenclatural codes. <em>EJT</em> is an archived and indexed journal that welcomes scientific contributions from all over the world, both in content and authorship. If you have any questions about <em>EJT</em>, please <a href="">contact us</a></p> EJT Consortium en-US European Journal of Taxonomy 2118-9773 <h3>Creative Commons Copyright Notices</h3> <div class="page"> <p>Authors who publish with this journal agree to the following terms:</p> <ol> <li class="show">Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a&nbsp;<a href="">Creative Commons Attribution License</a> (CC BY 4.0) that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.</li> <li class="show">Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.</li> <li class="show">Authors are NOT ALLOWED TO post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to taxonomic issues.</li> </ol> </div> A new genus and five new species of Neocoelidiinae leafhoppers from Brazil (Insecta: Hemiptera: Cicadellidae) with a key to males <p>A new Brazilian leafhopper genus of the subfamily Neocoelidiinae is proposed: <em>Takiyaella</em> gen.&nbsp;nov. based on <em>T</em>.&nbsp;<em>sexguttata</em> (Chiamolera &amp; Cavichioli, 2003) gen. et comb.&nbsp;nov. and five new species: <em>T</em>.&nbsp;<em>anomala</em> gen. et sp.&nbsp;nov., <em>T</em>.&nbsp;<em>coelhomarquesae</em> gen. et sp.&nbsp;nov., and <em>T</em>.&nbsp;<em>daniela</em> gen. et sp.&nbsp;nov. from the state of Rio de Janeiro; <em>T</em>.&nbsp;<em>cavichiolii</em> gen. et sp.&nbsp;nov. from the state of Paraná, and <em>T</em>.&nbsp;<em>mejdalanii</em> gen. et sp.&nbsp;nov. from the state of Minas Gerais. A distribution map and identification key to <em>Takiyaella</em> species are provided.</p> Clayton C. Gonçalves Alexandre C. Domahovski Copyright (c) 2021 Clayton C. Gonçalves, Alexandre C. Domahovski 2021-10-18 2021-10-18 775 34–61 34–61 10.5852/ejt.2021.775.1545 Two novel species of subgenus Russula crown clade (Russulales, Basidiomycota) from China <p>Two novel species of Russula (Russulaceae, Russulales), <em>R. coronaspora</em> Y.Song sp.&nbsp;nov. and <em>R. minor</em> Y.Song sp.&nbsp;nov. belonging to subgenus Russula crown clade, are described based on both morphological and phylogenetic evidence. In morphology, <em>R. coronaspora</em> sp.&nbsp;nov. is mainly characterized by its distinct spores ornamented with sparse, cylindrical and isolated spines, which resemble coronavirus, and gelatinized pileipellis with pileocystidia mostly septate and sometimes branched; <em>R. minor</em> sp.&nbsp;nov. possesses a very small basidiocarp with pileus less than 2.5 cm in diameter., small basidia, easily peeling and gelatinized pileipellis with slender terminal cells and abundant SV+ pileocystidia. Positions of the two new species in both phylogenetic trees based on ITS and 5-locus sequences (nLSU, mtSSU,<em> rpb1</em>, <em>rpb2</em> and <em>tef1</em>) confirm their distinct taxonomic status.</p> Yu Song Xiu-Chao Xie Bart Buyck Copyright (c) 2021 Yu Song, Xiu-Chao Xie, Bart Buyck 2021-10-15 2021-10-15 775 15 33 10.5852/ejt.2021.775.1543 Pectotibialis paghmanensis Tshernyshev gen. nov. – a new genus and species of soft-winged flower beetle (Coleoptera, Cleroidea, Malachiidae) from Afghanistan <p>A new genus with a new species of soft-winged flower beetle, <em>Pectotibialis paghmanensis</em> Tshernyshev gen. et sp. nov. are described from Afghanistan. The new genus can be distinguished from the congeners of the tribe Apalochrini by the dark pectination in the apices of tibiae in both sexes, and the anterior tibiae which are hollowed at distal half, flattened and rounded femora, bituberculate basal parts of head and pronotum, two lamellate metathoracic appendages, tarsal comb above second tarsomere of anterior legs, and serrate antennae in the male. Based on the metathoracic appendages and comb in anterior legs would attribute this new species to the new genus <em>Dromanthomorphus</em> Pic, 1921, but all the other above-mentioned characters define its independent status and the designation of a new genus; <em>Pectotibialis</em> Tshernyshev gen. nov. The external appearance, special male characters and genitalia of the type species of the new genus are illustrated, and a distribution map is provided. A key to the <em>Apalochrus</em>-section of the tribe Apalochrini is provided.</p> Sergei E. Tshernyshev Copyright (c) 2021 Sergei E. Tshernyshev 2021-10-14 2021-10-14 775 1 14 10.5852/ejt.2021.775.1539 Morphology and zoogeography of the burrower-like gammarid Gammarus koshovi (Bazikalova, 1946) (Crustacea, Amphipoda, Gammaridae) – An overlooked and poorly known species in the Siberian fauna <p>A comprehensive survey of several lakes in the Tuva Republic of Russia yielded, besides the ubiquitous <em>Gammarus lacustris</em>, the discovery of a small freshwater, lacustrine amphipod not previously recorded in this area. A comparative study of the Tore-Khol Lake gammarid population, probably conspecific with <em>Gammarus koshovi</em> (Bazikalova, 1946) originating from Khubsugul Lake, was conducted. The species, <em>G.&nbsp;koshov</em>i, is characterized by a specific habitus: a small-sized compact body, all limbs shortened, carpi of pereopods (PIII and PIV) reduced, coxal plates broad, and pereopod dactyli sturdy. It has been suggested that juveniles of the euryoecious <em>G.&nbsp;lacustris</em> or other large species could be confused with the relatively small <em>G.&nbsp;koshovi</em>. Consequently, we decided to present the distribution of gammarid species throughout south Siberia and Mongolia, referring to the sequences of works primarily by Soviet authors, which may be hard to access by international readers. We discuss affinity with related groups, distribution, and ecology of <em>G.&nbsp;koshovi</em> to better understand their evolution. Additionally, the zoobenthic species diversity of widely represented groups in the ecosystems of Tore-Khol Lake is briefly reviewed. An identification key for the Siberian <em>Gammarus</em> with 10 species is provided.</p> Dmitry Sidorov Elena Yalysheva Mariana Sharyi-Ool Copyright (c) 2021 Dmitry Sidorov, Elena Yalysheva, Mariana Sharyi-Ool 2021-10-13 2021-10-13 775 178 199 10.5852/ejt.2021.774.1541 The caridean shrimps of the genus Lysmata Risso, 1816 (Decapoda: Lysmatidae) from Madagascar collected during the Atimo-Vatae expedition: a new species and two new records <p>The present study focuses on shrimps belonging to the genus <em>Lysmata</em> Risso, 1816, collected from Madagascar during the Atimo Vatae expedition carried out in 2010. <em>Lysmata</em> <em>malagasy</em> sp. nov. is a new species belonging to the clade named “long accessory ramous” or “cosmopolitan” in previous phylogenetic studies. The new species can be distinguished from the only two other representatives of this group in the Indo-west Pacific, <em>L. ternatensis</em> De Man, 1902, and<em> L. trisetacea</em> (Heller, 1861), by the accessory ramus of the lateral antennular flagellum consisting of four elongated articles. <em>Lysmata lipkei</em> Okuno &amp; Fiedler, 2010 is reported here from Madagascar with a remarkable extension of its known range after its original description from Japan. This species has also been reported from Singapore and, as alien species, from Brazil. Lastly, <em>L. kuekenthali</em> De Man, 1902 known from numerous localities in the Indo-West Pacific biogeographic area, is reported for the first time from Madagascar. Results of the present morphological and molecular analyses suggest that <em>L. hochi</em> Baeza &amp; Anker, 2008 from the Caribbean Sea is a synonym of the Indo-West Pacific <em>L. kuekenthali</em>, and thus the latter species is alien in the western Atlantic.</p> Hossein Ashrafi J. Antonio Baeza Zdeněk Ďuriš Copyright (c) 2021 Hossein Ashrafi, J. Antonio Baeza, Zdeněk Ďuriš 2021-10-12 2021-10-12 775 155 177 10.5852/ejt.2021.774.1535 Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini) <p>Three new species of the genus <em>Guiodytes</em> Tian, 2013 are described from the limestone caves of Guangxi Zhuang Autonomous Region, southern China: <em>Guiodytes weii</em> Huang &amp; Faille sp.&nbsp;nov. and <em>Guiodytes yueliangensis </em>Huang &amp; Tian sp.&nbsp;nov. from Dapo Dong and Huang Dong caves, respectively, in Huanjiang County, northernmost Guangxi; <em>Guiodytes inexpectatus </em>Tian &amp; Zhou sp.&nbsp;nov. from the Zhuzhu Dong cave in Longzhou County, southwestern Guangxi. All of the six known species of <em>Guiodytes</em> are endemic to Guangxi, ranging from the southwest to the northernmost. A modified key to species and a distribution map for&nbsp;<em>Guiodytes</em> are provided.</p> Sunbin Huang Jiajun Zhou Mingyi Tian Arnaud Faille Copyright (c) 2021 Sunbin Huang, Jiajun Zhou, Mingyi Tian, Arnaud Faille 2021-10-11 2021-10-11 775 135 154 10.5852/ejt.2021.774.1537 Middle Ordovician Drepanoistodus (Vertebrata, Conodonta) from Baltica, with description of three new species <p><em>Drepanoistodus basiovalis</em> (Sergeeva, 1963) is a common conodont species in Middle Ordovician strata of Baltica. For many years it has been widely accepted that the species encompasses a wide range of morphological plasticity. Hence, several different morphotypes that significantly deviate from the holotype have nonetheless been included in the broad species concept. In this study, we performed a detailed taxonomical study on 112 predominantly well-preserved specimens (geniculate elements) from the St. Petersburg region of Russia; 37 of these were selected for morphometric analyses together with 21 well-illustrated specimens from the published literature. The results demonstrate that, among the morphotypes that share some characteristics with <em>D.&nbsp;basiovalis</em> sensu lato, at least five species can be readily distinguished. Hence, three new species – <em>Drepanoistodus iommii</em> sp.&nbsp;nov., <em>D.&nbsp;svendi</em> sp.&nbsp;nov. and <em>D.&nbsp;viirae</em> sp.&nbsp;nov. – are here added to the previously known <em>D.&nbsp;basiovalis</em> and <em>D.&nbsp;contractus</em> (Lindström, 1955). In addition, some specimens were left under open nomenclature and assigned to <em>Drepanoistodus</em> aff. <em>basiovalis</em> and <em>D</em>. cf.&nbsp;<em>suberectus</em> (Branson &amp; Mehl, 1933). In order to objectively compare the <em>Drepanoistodus</em> taxa and test the validity of the new species, we performed a Principal Component Analysis combined with non-parametric (PERMANOVA) tests based on 21 morphological characters.</p> Jan Audun Rasmussen Mats E. Eriksson Anders Lindskog Copyright (c) 2021 Jan Audun Rasmussen, Mats E. Eriksson, Anders Lindskog 2021-10-08 2021-10-08 775 106–134 106–134 10.5852/ejt.2021.774.1533 Five new and five known species of the genus Tylencholaimus de Man, 1876 (Nematoda: Dorylaimida: Tylencholaimoidea) from Western Ghats of India <p>Five new and five known species of the genus <em>Tylencholaimus</em> de Man, 1876 are described from the Western Ghats of India. <em>Tylencholaimus</em> <em>macroamphidius</em> sp. nov. has a 0.61–0.85 mm long body, angular lips, large amphid, 8.0–9.0&nbsp;μm long odontostyle, odontophore with asymmetrical basal knobs, pharyngeal bulb expanding gradually, female genital system mono-prodelphic, and a rounded to conoid tail. <em>Tylencholaimus</em> <em>shamimi</em> sp. nov. has a 0.57–0.71 mm long body, 6.0–7.0&nbsp;μm long odontostyle, odontophore with basal thickening, pharyngeal bulb expanding gradually, female genital system amphidelphic and a convex-conoid tail. <em>Tylencholaimus</em> <em>southindicus</em> sp. nov. has a 0.44–0.55 mm long body, lip region with distinct labial disc, 4.5–5.5&nbsp;μm long odontostyle, odontophore with minute basal knobs, pharyngeal expansion abrupt, female genital system mono-prodelphic, and a rounded-conoid to rounded tail. <em>Tylencholaimus</em> <em>striatus</em> sp. nov. has a 0.30–0.34 mm long body with distinctly striated cuticle, 5.0–5.5&nbsp;μm long odontostyle, odontophore with minute basal knobs, pharyngeal bulb expanding abruptly, female genital system mono-prodelphic, and a conoid tail with bluntly rounded terminus. <em>Tylencholaimus</em> <em>tamiliensis</em> sp. nov. has 0.51–0.58 mm long body, 5.5–6.0&nbsp;μm long odontostyle, odontophore with minute basal knobs, pharyngeal bulb expanding gradually, female genital system mono-prodelphic and tail rounded to conoid with sunken terminus. Five known species of the genus<em> Tylencholaimus </em>viz. <em>T.&nbsp;mirabilis</em>, <em>T</em>. <em>teres</em>, <em>T</em>. <em>micronanus</em>, <em>T</em>.<em> ibericus</em> and <em>T</em>. <em>cosmos</em> also recorded from the region and redescribed/illustrated.</p> Md Niraul Islam Wasim Ahmad Copyright (c) 2021 Md Niraul Islam, Wasim Ahmad 2021-10-07 2021-10-07 775 58–105 58–105 10.5852/ejt.2021.774.1531 Aretopsis sandybrucei, a new deep-water shrimp (Malacostraca: Decapoda: Caridea: Alpheidae) from the Coral Sea <p>A new alpheid shrimp, <em>Aretopsis sandybrucei</em> sp. nov., is described on the basis of three specimens collected from three deep-water banks in the Chesterfield Plateau of the Coral Sea, between New Caledonia and Queensland, Australia, at a depth range of 280–550 m. The new species is the first known deep-water species of the genus <em>Aretopsis</em> De Man, 1910, with its two congeners, <em>A. amabilis</em> De Man, 1910 and <em>A. manazuruensis</em> Suzuki, 1971, being confined to the lower intertidal and shallow subtidal areas, to a maximum depth of 50 m. Based on morphological grounds, <em>A. sandybrucei</em> sp. nov. can be easily separated from <em>A. amabilis</em> and <em>A. manazuruensis</em> by the generally much stouter third to fifth pereiopods, with their dactyli each bearing a very small secondary unguis on the flexor margin, in comparison to a much stronger one in the other two species. <em>Aretopsis sandybrucei</em> sp. nov. also differs from <em>A. amabilis</em> and <em>A. manazuruensis</em> by the less contrasting colour pattern, including the more translucent, dull yellowish chelae and tail fan. An association of <em>A. sandybrucei</em> sp. nov. with a deep-water hermit crab appears to be highly likely due to the presence of several large hermit crabs (Paguridae) in at least one of the three dredge/bottom trawl hauls, which was also containing a paratype of the new species. The taxonomic status of <em>A. amabilis</em> and <em>A. manazuruensis</em> is discussed.</p> Hossein Ashrafi Zdeněk Ďuriš Arthur Anker Copyright (c) 2021 Hossein Ashrafi, Zdeněk Ďuriš, Arthur Anker 2021-10-06 2021-10-06 775 42–57 42–57 10.5852/ejt.2021.774.1527 Four new genera of the Ceratonotus group (Copepoda, Harpacticoida), including the description of two new species <p>In the framework of an ongoing extensive phylogenetic evaluation of the <em>Ceratonotus</em> group (Copepoda, Harpacticoida, Cletodidae), <em>Poropsyllus menzelae</em> gen. et sp.&nbsp;nov. from the sublittoral of south-western Cyprus (eastern Mediterranean Sea) and <em>Paratouphapleura aaroni</em> gen. et sp.&nbsp;nov. from the western Weddell Sea (Antarctica) are described. Both new species fit the autapomorphies of the <em>Ceratonotus</em> group but cannot be assigned to any of the genera so far known. Instead, each new species presents a set of derived characters that justify their placement in new genera, <em>Poropsyllus</em> gen.&nbsp;nov. and <em>Paratouphapleura</em> gen.&nbsp;nov., respectively. Furthermore, a comparison of the species placed in <em>Ceratonotus</em> Sars revealed that because of exclusive morphological deviations, <em>Ceratonotus concavus</em> Conroy-Dalton, <em>C.&nbsp;steiningeri</em> George, <em>C.&nbsp;tauroides</em> George, and <em>C.&nbsp;vareschii</em> George should be excluded from <em>Ceratonotus</em> and transferred to a new monophylum, <em>Tauroceratus</em> gen.&nbsp;nov. Likewise, <em>Polyascophorus monoceratus</em> George, Wandeness &amp;#38; Santos is characterized by several apomorphies that justify its transfer from <em>Polyascophorus</em> to a new taxon, <em>Pseudopolyascophorus</em> gen.&nbsp;nov. The <em>Ceratonotus</em> group is therefore increased to 31&nbsp;species assigned to 13&nbsp;genera. The systematic modifications conducted and resulting phylogenetic consequences are discussed in detail.</p> Kai Horst GEORGE Copyright (c) 2021 Kai Horst GEORGE 2021-10-05 2021-10-05 775 1 41 10.5852/ejt.2021.774.1525