European Journal of Taxonomy <p>The <em>European Journal of Taxonomy</em> is a peer-reviewed international journal in descriptive taxonomy, covering the eukaryotic world. Its content is fully electronic and <a href="">Open Access</a>. It is published and funded by a <a href="">consortium</a> of European natural history institutions. Neither authors nor readers have to pay fees. All articles published in <em>EJT</em> are compliant with the different nomenclatural codes. <em>EJT</em> is an archived and indexed journal that welcomes scientific contributions from all over the world, both in content and authorship. If you have any questions about <em>EJT</em>, please <a href="">contact us</a></p> EJT Consortium en-US European Journal of Taxonomy 2118-9773 <h3>Creative Commons Copyright Notices</h3> <div class="page"> <p>Authors who publish with this journal agree to the following terms:</p> <ol> <li class="show">Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a&nbsp;<a href="">Creative Commons Attribution License</a> (CC BY 4.0) that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.</li> <li class="show">Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.</li> <li class="show">Authors are NOT ALLOWED TO post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to taxonomic issues.</li> </ol> </div> New species, revision, and phylogeny of Ronzotherium Aymard, 1854 (Perissodactyla, Rhinocerotidae) <p><em>Ronzotherium</em> is one of the earliest Rhinocerotidae in Europe, which first appeared just after the Eocene/Oligocene transition (Grande Coupure), and became extinct at the end of the Oligocene. It is a large-sized rhinocerotid, with a special position in the phylogeny of this group, as being one of the earliest-branching true Rhinocerotidae. However, its intra-generic systematics has never been tested through computational phylogenetic methods and it is basically unknown. Its taxonomical history has gone through numerous complications, and thus we aim to provide here a complete revision of this genus, through phylogenetic methods. After a re-examination of all type specimens (five supposed species) as well as of most well-preserved specimens from all over Europe and ranging through the complete Oligocene epoch, we performed a parsimony analysis to test the position of some problematic specimens. According to our results, five species can be distinguished, <em>Ronzotherium velaunum</em> (type species), <em>R. filholi</em>, <em>R. elongatum</em> and <em>R. romani</em> as well as a new species: <em>R. heissigi</em> sp. nov. We also drastically re-interpret its anatomy and show that the ‘short-limbed’ “<em>Diaceratherium</em>” <em>massiliae</em>, described from Southern France, can be considered as a junior synonym of <em>R. romani</em>. Finally, we exclude the Asian species “<em>Ronzotherium</em>” <em>orientale</em> and “<em>Ronzotherium</em>” <em>brevirostre</em> from <em>Ronzotherium</em> and we consider <em>R. kochi</em> as a junior synonym of <em>R. filholi</em>.</p> Jérémy Tissier Pierre-Olivier Antoine Damien Becker Copyright (c) 2021 Jérémy Tissier, Pierre-Olivier Antoine, Damien Becker 2021-06-14 2021-06-14 753 1 80 10.5852/ejt.2021.753.1389 Revision of the spider genus Stygopholcus (Araneae, Pholcidae), endemic to the Balkan Peninsula <p>The genus <em>Stygopholcus</em> Kratochvíl, 1932 is endemic to the Balkan Peninsula and includes only four nominal species: the epigean <em>S.&nbsp;photophilus</em> Senglet, 1971 in the south (Greece to Albania) and the ‘northern clade’ consisting of three troglophile species ranging from Croatia to Albania: <em>S.&nbsp;absoloni</em> (Kulczyński, 1914); <em>S.&nbsp;skotophilus</em> Kratochvíl, 1940; and<em> S.&nbsp;montenegrinus</em> Kratochvíl, 1940 (original rank re-established). We present redescriptions of all species, including extensive data on ultrastructure, linear morphometrics of large samples, and numerous new localities. We georeference previously published localities as far as possible, correct several published misidentifications, and clarify nomenclatorial problems regarding the authority of <em>Stygopholcus</em> and the identity of the type species <em>S.&nbsp;absoloni</em>. We suggest that the ‘northern clade’ has a relict distribution, resulting from past and present geologic and climatic factors. Future work on <em>Stygopholcus</em> should focus on the southern Dinarides, combining dense sampling with massive use of molecular data.</p> Bernhard A. Huber Martina Pavlek Marjan Komnenov Copyright (c) 2021 Bernhard A. Huber, Martina Pavlek, Marjan Komnenov 2021-06-11 2021-06-11 753 1–60 1–60 10.5852/ejt.2021.752.1391 Review of the millipede genus Levizonus Attems, 1898, with description of a new species from the Far East of Russia (Diplopoda, Polydesmida, Xystodesmidae) <p>The genus <em>Levizonus</em> Attems, 1898 is rediagnosed and shown to contain eight species from Russia (Far East), North Korea, Japan and North-East China. One species is described here as new to science: <em>Levizonus nakhodka</em> sp. nov. A new formal synonym is proposed: <em>Levizonus circularis</em> Takakuwa, 1942 = <em>Levizonus variabilis</em> Lokschina &amp; Golovatch, 1977 syn. nov., the valid name being the former. <em>Levizonus circularis</em> Takakuwa, 1942 is recorded for the fauna of China for the first time. All currently known species of <em>Levizonus</em> are included in a key, mapped and discussed.</p> Elena V. Mikhaljova Copyright (c) 2021 Elena V. Mikhaljova 2021-06-10 2021-06-10 753 159 184 10.5852/ejt.2021.751.1387 Three new genera and five new species of the tribe Meconematini (Orthoptera: Tettigoniidae: Meconematinae) from Southwestern China <p>This paper deals with the brachypterous Meconematini, including three new genera, <em>Acosmetides</em> gen.&nbsp;nov., <em>Neocyrtopsides</em> gen.&nbsp;nov. and <em>Macrocosmetura</em> gen.&nbsp;nov. Five new species are described: <em>Acosmetides peltates</em> gen. et sp.&nbsp;nov., <em>Acosmetides dilobosa</em> gen. et sp.&nbsp;nov., <em>Acosmetides platycerca</em> gen. et sp.&nbsp;nov., <em>Neocyrtopsides bispina</em> gen. et sp.&nbsp;nov. and <em>Macrocosmetura truncata</em> gen. et sp.&nbsp;nov. Two new combinations are proposed: <em>Acosmetides trigentis</em> (Wang, Bian &amp; Shi, 2016) gen. et comb. nov. and <em>Neocyrtopsides platycata</em> (Shi &amp; Zheng, 1994) gen. et comb. nov.</p> Yan-Lin Chang Tao Wang Fu-Ming Shi Copyright (c) 2021 Yan-Lin Chang, Tao Wang, Fu-Ming Shi 2021-06-08 2021-06-08 753 140 158 10.5852/ejt.2021.751.1385 Monsters in the dark: systematics and biogeography of the stygobitic genus Godzillius (Crustacea: Remipedia) from the Lucayan Archipelago <p>Remipedia is a stygobitic group commonly associated with coastal anchialine caves. This class consists of 12 genera, ten of which are found within the Lucayan Archipelago. Herein, we describe a new species within the genus <em>Godzillius </em>from Conch Sound Blue Hole, North Andros Island, Bahamas. <em>Godzillius louriei </em>sp. nov. is the third known remipede observed from a subseafloor marine cave, and the first from the Godzilliidae. Remipedes dwell within notoriously difficult to access cave habitats and thus integrative and comprehensive systematic studies at family or genus level are often absent in the literature. In this study, all species of <em>Godzillius </em>are compared using morphological and molecular approaches. Specifically, the feeding appendages of <em>G. louriei </em>sp. nov., <em>G. fuchsi </em>Gonzalez, Singpiel &amp; Schlagner, 2013 and <em>G. robustus </em>Schram, Yager &amp; Emerson, 1986 were examined using scanning electron microscopy (SEM). Species of <em>Godzillius </em>are identified based on the spines of maxilla 1 segment 4 and by the denticles on the lacinia mobilis of the left mandible. A molecular phylogeny using the mitochondrial 16S rRNA and nuclear histone 3 genes recovered <em>G. louriei </em>sp. nov. within the <em>Godzillius </em>clade and 16S genetic distances revealed a 13–15% difference between species of <em>Godzillius</em>.</p> Lauren Ballou Thomas M. Iliffe Brian Kakuk Brett C. Gonzalez Karen J. Osborn Katrine Worsaae Kenneth Meland Kenneth Broad Heather Bracken-Grissom Jørgen Olesen Copyright (c) 2021 Lauren Ballou, Thomas M. Iliffe, Brian Kakuk, Brett C. Gonzalez, Karen J. Osborn, Katrine Worsaae, Kenneth Meland, Kenneth Broad, Heather Bracken-Grissom, Jørgen Olesen 2021-06-07 2021-06-07 753 115 139 10.5852/ejt.2021.751.1383 One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations <p>One new genus, <em>Zagrotes</em> gen. nov., and 19 new species of ground spiders (Gnaphosidae) are described from Iran: <em>Berinda bifurcata</em> sp.&nbsp;nov. (♂, Bushehr, Khuzestan; southwestern and southern Iran), <em>Berinda hoerwegi</em> sp.&nbsp;nov. (♂♀, Fars, Ilam, Kermanshah, Kurdistan; western and southcentral Iran), <em>Berlandina artaxerxes</em> sp.&nbsp;nov. (♂&nbsp;Yazd; central Iran), <em>Cryptodrassus iranicus</em> sp. nov. (♂, Kermanshah; western Iran),&nbsp;<em>Drassodes persianus</em> sp. nov. (♀, Kermanshah, Sistan &amp; Baluchistan; western and southeastern Iran), <em>Echemus caspicus</em> sp.&nbsp;nov. (♀, Golestan; northern Iran), <em>Gnaphosa qamsarica</em> sp.&nbsp;nov. (♀, Isfahan; central Iran), <em>Haplodrassus medes</em> sp.&nbsp;nov. (♂, Fars; southcentral Iran), <em>Haplodrassus qashqai</em> sp.&nbsp;nov. (♂♀, Hormozgan, Khuzestan, Lorestan; southwestern to southern Iran), <em>Marinarozelotes achaemenes</em> sp. nov. (♀, Kohgiluyeh &amp; Boyer-Ahmad; southwestern Iran), <em>Marjanus isfahanicus</em> sp.&nbsp;nov. (♀, Isfahan; central Iran), <em>Nomisia ameretatae</em> sp.&nbsp;nov. (♂, Tehran; northern Iran), <em>Prodidomus inexpectatus</em> sp.&nbsp;nov. (♂, Hormozgan; southern Iran), <em>Scotophaeus anahita</em> sp.&nbsp;nov. (♀, Isfahan; central Iran), <em>Scotophaeus elburzensis</em> sp.&nbsp;nov. (♀, Tehran, Zanjan; northwestern and northern Iran), <em>Sosticus montanus</em> sp.&nbsp;nov. (♀, Ilam; western Iran), <em>Synaphosus martinezi</em> sp. nov. (♂♀, Kohgiluyeh &amp; Boyer-Ahmad; southwestern Iran), <em>Zagrotes apophysalis</em> sp. nov. (♂♀, Hormozgan, Kohgiluyeh &amp; Boyer-Ahmad; southwestern to southern Iran) and <em>Zelotes hyrcanus</em> sp.&nbsp;nov. (♀, Mazandaran; northern Iran). These are the first records of the genera <em>Berinda</em> Roewer, 1928, <em>Echemus</em> Simon, 1878 and <em>Marjanus</em> Chatzaki, 2018 in Iran. Additionally, the previously unknown female of <em>Callipelis deserticola</em> Zamani &amp; Marusik, 2017 is described and illustrated, and <em>Berlandina mesopotamica</em> Al-Khazali, 2020 is recorded in Iran for the first time. Furthermore, <em>Berinda idae</em> Lissner, 2016 syn.&nbsp;nov. (Greece, Cyprus) is synonymized with <em>Berinda infumatus</em> (O. Pickard-Cambridge, 1872) comb.&nbsp;nov. (ex. Heser Tuneva, 2004; Greece, Tanzania, Egypt, Israel, introduced to Japan).</p> Alireza Zamani Maria Chatzaki Sergei L. Esyunin Yuri M. Marusik Copyright (c) 2021 Alireza Zamani, Maria Chatzaki, Sergei L. Esyunin, Yuri M. Marusik 2021-06-04 2021-06-04 753 68 114 10.5852/ejt.2021.751.1381 A morphological and molecular review of the genus Goniurosaurus, including an identification key <p>The genus <em>Goniurosaurus</em> (tiger geckos) currently consists of 23 species distributed in China, Japan and Vietnam. Several species complexes and recent discoveries of cryptic species pose challenges to the species identification, which is crucial to effectively implement the recent listing of the species from China and Vietnam in CITES Appendix II and the species from Japan in CITES Appendix III. Based on the results of our field work in northern Vietnam and data compiled from literature, we herein provide a taxonomic review of the genus <em>Goniurosaurus</em>. Our phylogenetic analyses showed that all recorded populations of tiger geckos from Vietnam, which were found to be monophyletic with low intra-specific genetic divergences, are assigned to one of the four species: <em>G. catbaensis</em>, <em>G. huuliensis</em>, <em>G.&nbsp;lichtenfelderi</em> or <em>G. luii</em>. Both genetic and morphological analyses confirm that the species from China and Vietnam can be split into three major groups. Based on the newly collected data, we provide an extended morphological description of the Vietnamese species. In addition, we provide an identification key for all <em>Goniurosaurus</em> species from China, Japan and Vietnam in order to assist authorities in the enforcement of the recent CITES listing.</p> Hai Ngoc Ngo Huy Quoc Nguyen Hieu Minh Tran Hanh Thi Ngo Minh Duc Le Laurenz Rafael Gewiss Mona van Schingen-Khan Truong Quang Nguyen Thomas Ziegler Copyright (c) 2021 Hai Ngoc Ngo, Huy Quoc Nguyen, Hieu Minh Tran, Hanh Thi Ngo, Minh Duc Le, Laurenz Rafael Gewiss, Mona van Schingen-Khan, Truong Quang Nguyen, Thomas Ziegler 2021-05-31 2021-05-31 753 38–67 38–67 10.5852/ejt.2021.751.1379 The pseudodichotomous Dasya sylviae sp. nov. (Delesseriaceae, Ceramiales) from 60–90 m mesophotic reefs off Bermuda <p>The red alga <em>Dasya sylviae</em> C.W.Schneid., M.M.Cassidy &amp; G.W.Saunders sp. nov. is described from mesophotic depths of 60–90 m off Bermuda. Genetic sequences (COI-5P, rbcL) and morphological characteristics show that this species is distinct from other known pseudodichotomous species of <em>Dasya</em>. Of ten current species in the genus reported from Bermuda, only three, <em>D.&nbsp;collinsiana</em> M.Howe, <em>D.&nbsp;cryptica</em> C.W.Schneid., Quach &amp; C.E.Lane and <em>D.&nbsp;punicea</em> (Zanardini) Menegh., share the overall pattern of pseudodichotomous branching in their axes; however, key morphological features easily distinguish them from<em> D.&nbsp;sylviae</em> sp.&nbsp;nov. The species most similar in habit to <em>D.&nbsp;sylviae</em> sp.&nbsp;nov. is <em>D.&nbsp;crouaniana</em> J.Agardh (type locality West Indies), but it bears shorter pseudolateral branches, and broader and longer tetrasporangial stichidia than the new species. Unique among the species of <em>Dasya</em>, <em>D.&nbsp;sylviae</em> sp.&nbsp;nov. lacks post-sporangial cover cells in tetrasporangial stichidia.</p> Craig W. Schneider Margaret M. Cassidy Gary W. Saunders Copyright (c) 2021 Craig W. Schneider, Margaret M. Cassidy, Gary W. Saunders 2021-05-27 2021-05-27 753 24 37 10.5852/ejt.2021.751.1377 Taxonomy of the New World bee genus Agapostemon Guérin-Méneville – new names and synonymies (Hymenoptera: Halictidae) <p>Many early taxonomic works on North American bees were published by Europeans using specimens collected in the New World, some with type locations so imprecise that uncertainty on the nomenclatural status remains to this day. Two examples come from Fabricius (1745–1808) who described <em>Andrena virescens</em> Fabricius, 1775 and <em>Apis viridula</em> Fabricius, 1793 from “America” and “Boreal America”, respectively. The former species of <em>Agapostemon</em> Guérin-Méneville, 1844 occurs across most of the United States and southern Canada, the latter presumed an endemic to Cuba. The type materials of these two taxa have never been compared to each other, though a morphology-based phylogenetic analysis placed both in distinct species groups. Here we synonymize <em>Apis viridula</em> under <em>Ag.&nbsp;virescens</em>, thereby making <em>Ag. femoralis</em> (Guérin-Méneville, 1844) available as the name for the Cuban species. A lectotype for <em>Ag. femoralis</em> (the type species for the genus <em>Agapostemon</em>) is hereby designated to stabilize this taxonomy. We also synonymize <em>Ag. obscuratus</em> Cresson, 1869 under <em>Ag.&nbsp;femoralis</em>, suggesting that it represents a dark colour polymorphism. As <em>Ag. cubensis</em> Roberts, 1972 is a junior secondary homonym of <em>Ag. cubensis</em> (Spinola, 1851), we offer <em>Ag. robertsi</em> as a replacement name for the former.</p> Cory S. Sheffield Lars Vilhelmsen Frederique Bakker Copyright (c) 2021 Cory S. Sheffield, Lars Vilhelmsen, Frederique Bakker 2021-05-25 2021-05-25 753 1–23 1–23 10.5852/ejt.2021.751.1375 The Smicridea (Smicridea) fasciatella species group (Trichoptera: Hydropsychidae) in Brazil: six new species and new distributional records <p>The <em>Smicridea</em> (<em>Smicridea</em>) <em>fasciatella</em> species group occurs from the southwestern USA, throughout Central America, the Greater Antilles islands, and most of South America, except for the Chilean subregion. It is characterized by the phallic apparatus being a simple tube with eversible internal sclerites at the apex. The <em>fasciatella</em> group is composed of 61 species, of which only 11 occur in Brazil, mainly in the Atlantic Forest biome in the southeastern region. In order to reduce the Linnean and Wallacean shortfalls for the Smicridea Brazilian fauna, we diagnose, describe, and illustrate males of six new species in the fasciatella group: <em>Smicridea</em> (<em>Smicridea</em>) <em>blahniki</em> Desiderio, Pes&nbsp;&amp; Hamada sp.&nbsp;nov.,<em> S.</em>&nbsp;(<em>Smicridea</em>) <em>brevitruncata</em> Desiderio, Pes &amp; Hamada sp. nov.,<em> S.&nbsp;</em>(<em>Smicridea</em>) <em>caaguara</em> Desiderio, Pes &amp; Hamada sp. nov., <em>S.</em>&nbsp;(<em>Smicridea</em>) <em>ipiranga</em> Desiderio, Pes &amp; Hamada sp. nov., <em>S.&nbsp;</em>(<em>Smicridea</em>) <em>jeaneae</em> Desiderio, Pes &amp; Hamada sp. nov., and <em>S.</em>&nbsp;(<em>Smicridea</em>) <em>polyacantha</em> Desiderio, Pes &amp;; Hamada sp. nov. Additionally, we provide distributional data for <em>S.&nbsp;</em>(<em>Smicridea</em>) <em>albosignata</em> Ulmer, 1907,<em> S.</em>&nbsp;(<em>Smicridea</em>) <em>bivittata</em> (Hagen, 1861), <em>S.</em>&nbsp;(<em>Smicridea</em>) <em>erecta</em> Flint, 1974, <em>S.&nbsp;</em>(<em>Smicridea</em>) <em>obliqua</em> Flint, 1974, <em>S.</em>&nbsp;(<em>Smicridea</em>) <em>paranensis</em> Flint, 1983, and<em> S.</em>&nbsp;(<em>Smicridea</em>) <em>sattleri</em> Denning &amp; Sykora, 1968. The number of<em> S.</em>&nbsp;(<em>Smicridea</em>) species in Brazil increases from 21 to 27 and <em>Smicridea</em> is recorded from the states of Acre, Amapá, and Sergipe for the first time.</p> Gleison Robson Desiderio Ana Maria Pes Vanderly Andrade-Souza Neusa Hamada Copyright (c) 2021 Gleison Robson Desiderio, Ana Maria Pes, Vanderly Andrade-Souza, Neusa Hamada 2021-05-24 2021-05-24 753 156 196 10.5852/ejt.2021.750.1371