European Journal of Taxonomy https://europeanjournaloftaxonomy.eu/index.php/ejt <p>The <em>European Journal of Taxonomy</em> is a peer-reviewed international journal in descriptive taxonomy, covering the eukaryotic world. Its content is fully electronic and <a href="https://europeanjournaloftaxonomy.eu/index.php/ejt/openaccess">Open Access</a>. It is published and funded by a <a href="https://europeanjournaloftaxonomy.eu/index.php/ejt/about">consortium</a> of European natural history institutions. Neither authors nor readers have to pay fees. All articles published in <em>EJT</em> are compliant with the different nomenclatural codes. <em>EJT</em> is an archived and indexed journal that welcomes scientific contributions from all over the world, both in content and authorship. If you have any questions about <em>EJT</em>, please <a href="mailto:ejteditorialoffice@gmail.com">contact us</a></p> EJT Consortium en-US European Journal of Taxonomy 2118-9773 <h3>Creative Commons Copyright Notices</h3> <div class="page"> <p>Authors who publish with this journal agree to the following terms:</p> <ol> <li class="show">Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a&nbsp;<a href="https://creativecommons.org/licenses/by/4.0/">Creative Commons Attribution License</a> (CC BY 4.0) that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.</li> <li class="show">Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.</li> <li class="show">Authors are NOT ALLOWED TO post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to taxonomic issues.</li> </ol> </div> The nautiloids from the Early Carboniferous Dalle à Merocanites of Timimoun, western Algeria https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1635 <p>Early Carboniferous coiled nautiloids from North Africa are virtually unknown. An assemblage of nine species, all from the family Trigonoceratidae, from the Dalle à <em>Merocanites</em> (Tournaisian-Viséan boundary interval) of Timimoun in western Algeria is described, being the most diverse Carboniferous nautiloid assemblage known from North Africa but much less diverse than the time-equivalent assemblages from Belgium and Ireland. The assemblage consists of the species <em>Maccoyoceras pentagonum</em> sp. nov., <em>Lispoceras orbis</em> sp. nov., <em>Thrincoceras devolvere</em> sp. nov., <em>Rineceras multituberculatum</em> sp. nov., <em>Rineceras rectangulatum</em> sp. nov., <em>Vestinautilus padus</em> sp. nov., <em>Vestinautilus concinnus</em> sp. nov., <em>Planetoceras destrictum</em> sp. nov. and <em>Planetoceras transforme</em> sp. nov. A morphometric analysis of <em>Maccoyoceras pentagonum</em> sp. nov. and <em>Lispoceras orbis</em> sp. nov. shows that the intraspecific variation in these species is within rather narrow limits.</p> Dieter Korn Luyi Miao Jürgen Bockwinkel Copyright (c) 2022 Dieter Korn, Luyi Miao, Jürgen Bockwinkel https://creativecommons.org/licenses/by/4.0 2022-01-28 2022-01-28 789 104 129 10.5852/ejt.2022.789.1635 The genus Alexeter Förster (Hymenoptera, Ichneumonidae) in China, with descriptions of three new species https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1633 <p>Three new species of <em>Alexeter</em> Förster, 1869 are reported from China,<em> A. flavicoxa</em> Li &amp; Sun sp. nov. collected from Sichuan, <em>A. flaviventris</em> Li &amp; Sun sp. nov. from Beijing, Guizhou and <em>A. melanicus</em> Li &amp; Sun sp. nov. from Guizhou. A key to species of <em>Alexeter</em> known in China is provided.</p> Tao Li Shu-Ping Sun Copyright (c) 2022 Tao Li, Shu-Ping Sun https://creativecommons.org/licenses/by/4.0 2022-01-26 2022-01-26 789 81 103 10.5852/ejt.2022.789.1633 Six new species of Amblypsilopus Bigot (Diptera: Dolichopodidae: Sciapodinae) and a key to species of the Afrotropical mainland https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1631 <p>Six new species of <em>Amblypsilopus </em>Bigot, 1888 from the Afrotropical region are described and illustrated: <em>A.&nbsp;gabonensis </em>sp.&nbsp;nov. from Gabon, <em>A.&nbsp;martini</em> sp.&nbsp;nov., <em>A.&nbsp;mufindiensis </em>sp.&nbsp;nov. and <em>A.&nbsp;udzungwensis </em>sp.&nbsp;nov. from Tanzania,<em> A.&nbsp;milleri </em>sp.&nbsp;nov. from South Africa, and <em>A.&nbsp;spiniscapus </em>sp.&nbsp;nov. from Ivory Coast. <em>Amblypsilopus gabonensis </em>sp.&nbsp;nov. and <em>A.&nbsp;spiniscapus </em>sp.&nbsp;nov. are remarkable in having a setose antennal scape. <em>Amblypsilopus martini</em> sp.&nbsp;nov. and <em>A.&nbsp;mufindiensis </em>sp.&nbsp;nov. are peculiar in bearing very long setae on the antennal pedicel, mesonotum and scutellum. <em>Amblypsilopus milleri </em>sp.&nbsp;nov. is considered a member of the South African <em>A.&nbsp;fasciatus </em>species group despite its unmodified wing venation. The new species differ also from other similar representatives of the genus in the morphology of the male cercus and the male secondary sexual characters on the legs. The male of <em>A.&nbsp;bevisi </em>(Curran, 1927) is described for the first time. The following species described from females are considered doubtful (nomina dubia): <em>A.&nbsp;flavicollis</em> (Becker, 1923), <em>A.&nbsp;rectangularis</em> (Parent, 1937), <em>A.&nbsp;sudanensis</em> (Parent, 1939) and <em>A.&nbsp;tropicalis</em> (Parent, 1933).<em> Amblypsilopus subfascipennis</em> (Curran, 1926) has a strongly sinuate wing vein dm-m and is transferred to <em>Chrysosoma</em> Guérin-Méneville, 1831 (comb. nov.). <em>Chrysosoma ungulatum </em>Parent, 1941 from Príncipe with an almost straight vein dm-m is transferred to<em> Amblypsilopus</em> (comb. nov.). <em>Chrysosoma centrale </em>Becker, 1923 from Tanzania also has a straight wing vein dm-m and modified fore tarsus, being very close in habitus to <em>A.&nbsp;steelei</em> Grichanov, 1996, and is also transferred to<em> Amblypsilopus</em> (comb. nov.).<em> Chrysosoma asperum </em>Parent, 1933 from South Africa has the same characters, is almost identical to South African <em>A.&nbsp;bevisi </em>and is also transferred here to<em> Amblypsilopus</em> (comb. nov.) and newly synonymized with <em>A.&nbsp;bevisi </em>(syn. nov.). Presently, 66 reliable species of <em>Amblypsilopus </em>are known from the Afrotropical region, including 37 from the Afrotropical mainland. A revised identification key to males of species of the Afrotropical mainland is provided. New records are given for some known species. The species diversity of the Afrotropical fauna and variability of morphological characters in the genus <em>Amblypsilopus </em>are briefly discussed.</p> Igor Ya. Grichanov Copyright (c) 2022 Igor Ya. Grichanov https://creativecommons.org/licenses/by/4.0 2022-01-25 2022-01-25 789 49–80 49–80 10.5852/ejt.2022.789.1631 Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1627 <p>The diversity of Palpigradi is not evenly distributed among its different branches. The widespread genus <em>Eukoenenia</em> includes 80% of the species, while the other genera are poorly known. <em>Allokoenenia</em> certainly is the most understudied genus because it is represented only by the African species <em>Allokoenenia afra</em> Silvestri, 1913. Its description is short and does not include many features depicted in modern taxonomy of Palpigradi. In this paper, we describe two troglobitic species of <em>Allokoenenia</em>, report the occurrence of a third species represented by an immature specimen from Brazilian caves, and provide brief notes on the morphology of <em>A.&nbsp;afra</em>. <em>Allokoenenia canhembora</em> sp.&nbsp;nov., <em>A. stygia</em> sp.&nbsp;nov., and <em>Allokoenenia</em> sp. differ from <em>A.&nbsp;afra</em> by several morphological features, including more elongated appendages and a greater number of blades on lateral organs. Thus, they are considered troglomorphic. These new species are vulnerable to extinction because they are endemic to a single or few caves directly impacted by mining activities and groundwater exploitation. This study represents the first step for the conservation of these species and their habitats, since Brazilian caves with rare troglobites cannot be irreversibly impacted. Also, it brings important contributions on the distribution and morphology of this enigmatic genus.</p> Maysa Fernanda Villela Rezende Souza Rodrigo Lopes Ferreira Copyright (c) 2022 Maysa Fernanda Villela Rezende Souza, Rodrigo Lopes Ferreira https://creativecommons.org/licenses/by/4.0 2022-01-24 2022-01-24 789 11–48 11–48 10.5852/ejt.2022.789.1627 Three new species of Oxyethira Eaton, 1873 (Trichoptera: Hydroptilidae) from New Caledonia https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1623 <p><em>Oxyethira</em> Eaton, 1873 is one of the most diverse genera of Hydroptilidae, comprising over 240 species distributed in all biogeographical regions. Here three new species of <em>Oxyethira </em>(<em>Trichoglene</em>) Neboiss, 1977 are described and illustrated from male specimens collected in New Caledonia<em>: O.&nbsp;</em>(<em>Trichoglene</em>)<em> hamus</em> sp.&nbsp;nov., recognized by the hook-shaped apex of the long inferior appendages in lateral view and by the posterior margin of segment IX with a trilobed appearance in ventral view; <em>O.&nbsp;</em>(<em>Trichoglene</em>)<em> rectangulata</em> sp.&nbsp;nov., recognized by the rectangular shape of the inferior appendages, which are totally fused and with two pairs of small setae on the inner face; and <em>O. </em>(<em>Trichoglene</em>)<em> spiralis </em>sp.&nbsp;nov., recognized by the strongly curvilinear shape of the subgenital process in dorsal and lateral views and by the long process spiralling around the ejaculatory duct at the phallus apex.</p> Bruna Maria Silva Cavalcante Kjell Arne Johanson Copyright (c) 2022 Bruna Maria Silva Cavalcante, Kjell Arne Johanson https://creativecommons.org/licenses/by/4.0 2022-01-12 2022-01-12 789 1–10 1–10 10.5852/ejt.2022.789.1623 Pseudocetherinae (Hemiptera: Reduviidae) revisited: phylogeny and taxonomy of the lobe-headed bugs https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1625 <p>The concept of the previously monogeneric subfamily Pseudocetherinae (Hemiptera: Heteroptera: Reduviidae) is revised and expanded. We here transfer <em>Gerbelius</em> Distant, 1903, <em>Kayanocoris</em> Miller, 1954, <em>Microvarus</em> Jeannel, 1917, <em>Paragerbelius</em> Miller, 1958, and <em>Voconia</em> Stål, 1866 from Reduviinae to Pseudocetherinae and treat <em>Kayanocoris</em>, <em>Microvarus</em>, <em>Paragerbelius</em>, and <em>Pseudocethera</em> Villiers, 1963 as junior synonyms of <em>Voconia</em>, resulting in new combinations for <em>Voconia conradti</em> (Jeannel, 1917) comb.&nbsp;nov., <em>V.&nbsp;ifana</em> (Villiers, 1963) comb.&nbsp;nov., <em>V.&nbsp;monodi</em> (Villiers, 1963) comb.&nbsp;nov., <em>V.&nbsp;motoensis</em> (Schouteden, 1929) comb.&nbsp;nov., <em>V.&nbsp;ornata</em> (Distant, 1903) comb.&nbsp;nov., <em>V.&nbsp;schoutedeni</em> (Villiers, 1964) comb.&nbsp;nov., <em>V.&nbsp;typica</em> (Miller, 1958) comb.&nbsp;nov., and <em>V.&nbsp;wegneri</em> (Miller, 1954) comb.&nbsp;nov.&nbsp;We also describe 23 new species of <em>Voconia</em>: <em>V.&nbsp;bakeri</em> sp.&nbsp;nov., <em>V.&nbsp;bracata</em> sp.&nbsp;nov., <em>V.&nbsp;brachycephala</em> sp.&nbsp;nov., <em>V.&nbsp;chrysoptera</em> sp.&nbsp;nov., <em>V.&nbsp;coronata</em> sp.&nbsp;nov., <em>V.&nbsp;decorata</em> sp.&nbsp;nov., <em>V.&nbsp;dolichocephala</em> sp.&nbsp;nov., <em>V.&nbsp;fasciata</em> sp.&nbsp;nov., <em>V.&nbsp;grandioculata</em> sp.&nbsp;nov., <em>V.&nbsp;hemera</em> sp.&nbsp;nov., <em>V.&nbsp;isosceles</em> sp.&nbsp;nov., <em>V.&nbsp;laosensis</em> sp.&nbsp;nov., <em>V.&nbsp;lasiosoma</em> sp.&nbsp;nov., <em>V.&nbsp;lirophleps</em> sp.&nbsp;nov., <em>V.&nbsp;loki</em> sp.&nbsp;nov., <em>V.&nbsp;mexicana</em> sp.&nbsp;nov., <em>V.&nbsp;minima</em> sp.&nbsp;nov., <em>V.&nbsp;nyx</em> sp.&nbsp;nov., <em>V.&nbsp;smithae</em> sp.&nbsp;nov., <em>V.&nbsp;tridens</em> sp.&nbsp;nov., <em>V.&nbsp;trinidadensis</em> sp.&nbsp;nov., <em>V.&nbsp;tuberculata</em> sp.&nbsp;nov., and <em>V.&nbsp;vittata</em> sp.&nbsp;nov.&nbsp;Lectotypes are designated for <em>Gerbelius confluens</em> Distant, 1903, <em>G</em><em>. typicus</em> Distant, 1903, <em>V</em><em>. conradti</em> comb.&nbsp;nov., <em>V.&nbsp;ornata</em> comb.&nbsp;nov., and <em>V.&nbsp;pallidipes</em> Stål, 1866. A revised diagnosis and description of Pseudocetherinae are provided along with photographs of the species and of the male genitalia of 13 pseudocetherine and five closely related reduviine species. An identification key to the two genera of Pseudocetherinae as well as a key to species of <em>Voconia</em> are presented. A phylogenetic hypothesis is proposed for the relationships of Pseudocetherinae using parsimony analyses of 77 morphological characters.</p> Stephanie Castillo Dávid Rédei Christiane Weirauch Copyright (c) 2022 Stephanie Castillo, Dávid Rédei, Christiane Weirauch https://creativecommons.org/licenses/by/4.0 2022-01-11 2022-01-11 789 1–95 1–95 10.5852/ejt.2022.788.1625 Two new species of Metacyclops Kiefer, 1927 (Copepoda, Cyclopoida) from Thailand and an up-to-date key to the species recorded in Asia https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1621 <p><em>Metacyclops sakaeratensis</em> sp.&nbsp;nov. and <em>M.&nbsp;brancelji</em> sp.&nbsp;nov. are described as being present in the mountainous areas of Northeastern and Southern Thailand, respectively. Two new species resemble <em>M.&nbsp;woni</em> in both sexes, but they are easily distinguished from their Cambodian relative by having: 1)&nbsp;transverse suture on the dorsal surface of the genital double-somite, 2) serrated hyaline frill on the posterior margin of third pedigerous somite, 3) different length / width ratio of caudal ramus, and 4) a row of spinule on caudal surface of intercoxal sclerite of the third swimming legs. The significant differences between<em> M. sakaeratensis</em> sp.&nbsp;nov. and <em>M.&nbsp;brancelji</em> sp.&nbsp;nov. are present in both sexes, including the body size, integumental ornamentation of the body surface, length / width ratio of caudal ramus, armature of the fifth swimming leg, and the male sixth swimming leg. In addition, an up-to-date key to the female of all fifteen species of <em>Metacyclops</em> recorded in Asia is provided.</p> Sujeephon Athibai Koraon Wongkamhaeng Chaichat Boonyanusith Copyright (c) 2022 Sujeephon Athibai, Koraon Wongkamhaeng, Chaichat Boonyanusith https://creativecommons.org/licenses/by/4.0 2022-01-10 2022-01-10 789 146 181 10.5852/ejt.2021.787.1621 Reinstatement of Perinereis bairdii (Webster, 1884) and description of P. websteri sp. nov. from Bermuda, including the reproductive morphology of two Atlantic Perinereis species (Annelida: Errantia: Phyllodocida) https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1619 <p><em>Perinereis anderssoni</em> Kinberg, 1865 originally described from Brazil has been attributed with a wide distribution on the American Atlantic coasts, ranging from Brazil to Bermuda, due to the synonymy of <em>Nereis bairdii</em> Webster, 1884 and <em>Nereis (Perinereis) melanocephala</em> McIntosh, 1885. In this paper, the synonymy of <em>Nereis bairdii</em> with <em>Perinereis anderssoni</em> is rejected based on a re-examination of the syntypes of <em>N. bairdii</em> which are found to contain two species requiring designation of a lectotype and paralectotypes here redescribed as <em>Perinereis bairdii</em> (Webster, 1884) and the remaining specimens are here described as <em>Perinereis websteri</em> sp.&nbsp;nov. The synonymy of <em>Nereis</em> (<em>Perinereis</em>) <em>melanocephala</em> with <em>P.&nbsp;anderssoni</em> is rejected and the synonymy with <em>P.&nbsp;bairdii</em> proposed by previous authors is accepted here. A description of <em>P.&nbsp;anderssoni</em>, a redescription of <em>P.&nbsp;floridana</em>, and a description of the males of <em>P.&nbsp;cariboea</em> with additional material are provided for comparison. A key to identifying all Atlantic <em>Perinereis</em> species is also included.</p> Víctor Manuel Conde-Vela Copyright (c) 2022 Víctor Manuel Conde-Vela https://creativecommons.org/licenses/by/4.0 2022-01-06 2022-01-06 789 104–145 104–145 10.5852/ejt.2021.787.1619 On Thaicypris panhai gen. et sp. nov., a new herpetocypridine ostracod (Crustacea: Ostracoda: Cyprididae) from Thailand https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1617 <p>A new genus, <em>Thaicypris</em> gen.&nbsp;nov., in the tribe Herpetocypridini Kaufmann, 1900 of the subfamily Herpetocypridinae Kaufmann, 1900 is established to accommodate a new species from Thailand. The present contribution deals with the description of a new genus and species, <em>Thaicypris panhai</em> gen. et sp. nov., which is mainly characterized by the distinctive and raised, inwardly displaced selvage at the postero-ventral part of the right valve (RV) that is not parallel to the valve margin, the absence of an anterior inner list on the RV, the prominent and elevated double inner list on the posterior part of the left valve, the small and three-segmented Rome organ on the first antenna (A1), the spatulated terminal segment of the maxillular (Mx1) palp, the slender caudal ramus (CR) with long and thin Sp&nbsp;seta, the presence of basal triangle on the CR attachment, and the pointed projection at the terminal segment base of the prehensile palps. The hemipenis of the new genus and species is outstanding, especially the medial lateral shield which has a long, beak-shaped protrusion on the distal part. The discovery of this Thai taxon is the first record of the tribe Herpetocypridini in Thailand and the second species of the tribe in Southeast Asia.</p> Sukonthip Savatenalinton Copyright (c) 2022 Sukonthip Savatenalinton  https://creativecommons.org/licenses/by/4.0 2022-01-05 2022-01-05 789 86–103 86–103 10.5852/ejt.2021.787.1617 Two new species of Ageniella Banks, 1912 (Hymenoptera: Pompilidae) from Brazil and updated keys https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1615 <p><em>Ageniella</em> Banks, 1912 is a paraphyletic group exclusive to the Nearctic and Neotropical regions. The genus has a remarkable morphological diversity among species and strong sexual dimorphism, hampering taxonomic studies of it. Herein, we add two new species to the Neotropical fauna of <em>Ageniella</em>: <em>A.&nbsp;caerulea</em> sp.&nbsp;nov. belonging to the subgenus <em>Ameragenia </em>and <em>A.&nbsp;ruschi</em> sp.&nbsp;nov. belonging to the subgenus <em>Priophanes</em>; both species are described and illustrated. A brief discussion of subgeneric characters and an updated taxonomic key to the species of <em>Ageniella </em>known from Brazil are provided.</p> Márcia Rapoza Cecilia Waichert Copyright (c) 2022 Márcia Rapoza, Cecilia Waichert https://creativecommons.org/licenses/by/4.0 2022-01-04 2022-01-04 789 71–85 71–85 10.5852/ejt.2021.787.1615