European Journal of Taxonomy <p>The <em>European Journal of Taxonomy</em> is a peer-reviewed international journal in descriptive taxonomy, covering the eukaryotic world. Its content is fully electronic and <a href="">Open Access</a>. It is published and funded by a <a href="">consortium</a> of European natural history institutions. Neither authors nor readers have to pay fees. All articles published in <em>EJT</em> are compliant with the different nomenclatural codes. <em>EJT</em> is an archived and indexed journal that welcomes scientific contributions from all over the world, both in content and authorship. If you have any questions about <em>EJT</em>, please <a href="">contact us</a></p> EJT Consortium en-US European Journal of Taxonomy 2118-9773 <h3>Creative Commons Copyright Notices</h3> <div class="page"> <p>Authors who publish with this journal agree to the following terms:</p> <ol> <li class="show">Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a&nbsp;<a href="">Creative Commons Attribution License</a> (CC BY 4.0) that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.</li> <li class="show">Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.</li> <li class="show">Authors are NOT ALLOWED TO post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to taxonomic issues.</li> </ol> </div> Checklist and a new species of Lippia (Verbenaceae) from the Diamantina Plateau, Minas Gerais, Brazil <p>The Diamantina Plateau is located in the central region of the Espinhaço Range, in the State of Minas Gerais, which is dominated by campo rupestre formations. We describe a new species of <em>Lippia&nbsp;</em>L., endemic to the Diamantina Plateau, and provide an annotated checklist and identification key for the 17 species of the genus occurring in the area. <em>Lippia raoniana</em> P.H.Cardoso &amp; Salimena sp. nov. is mainly distinguished by its ovate leaves with adaxial and abaxial surfaces densely covered by sessile glandular trichomes, and drupaceous fruits with two pyrenes. It is known only from two populations, and thus can be provisionally considered as Critically Endangered. Details on the species’ ecology, as well as a comparison with morphologically similar species, a distribution map, and field photographs, are provided. In this work, we also present pictures of the 17 species in their habitats, and we hope that these illustrations will help in the identification and conservation of these taxa.</p> Pedro Henrique Cardoso Luiz Menini Neto Marcelo Trovó Fátima Regina Gonçalves Salimena Copyright (c) 2021 Pedro Henrique Cardoso, Luiz Menini Neto, Marcelo Trovó, Fátima Regina Gonçalves Salimena 2021-01-28 2021-01-28 733 42 55 10.5852/ejt.2021.733.1219 Two new species of Cypricercinae McKenzie, 1971 (Crustacea: Ostracoda) from Thailand <p>Two new species of the subfamily Cypricercinae McKenzie, 1971 are described from the Western part of Thailand: <em>Pseudostrandesia ratchaburiensis</em> sp.&nbsp;nov. and <em>Strandesia prachuapensis</em> sp.&nbsp;nov. <em>Pseudostrandesia ratchaburiensis</em> sp.&nbsp;nov. is mainly characterized by a flange on the antero-ventral part of the left valve (LV), a markedly large β seta on the mandibular (Md) palp, serrated bristles on the third endite of the maxillula (Mx1), a slender caudal ramus (CR) with a long claw Ga (length ca half that of the ramus) and a relatively low number (13) of spiny whorls in the Zenker’s organ. The discovery of both males and females of Pseudostrandesia ratchaburiensis sp.&nbsp;nov. in the present study constitutes the first report of a sexual population in this genus, thereby allowing for a comparison of the male reproductive organs (hemipenis and Zenker’s organ) from a new species with those of other genera of Cypricercinae. <em>Strandesia prachuapensis</em> sp.&nbsp;nov. is most closely related to <em>Strandesia odiosa</em> (Moniez, 1892) and <em>Strandesia flavescens</em> Klie, 1932 as they bear similar anterior flanges on the right valve (RV). The key diagnostic features of the new <em>Strandesia</em> species are a large carapace (ca 1.5 mm), an angulated antero-ventral part of the LV, a weak and small anterior inner list on the LV, an anterior flange on the RV, a markedly small aesthetasc Y on the second antenna, a large β seta on the Md-palp, smooth bristles on the third endite of the Mx1 and a slender CR with a short claw Ga (length ca ⅓ of the ramus). In addition, <em>Pseudostrandesia complexa</em> (Victor &amp; Fernando, 1981) comb. nov. is here proposed.</p> Sukonthip Savatenalinton Copyright (c) 2021 Sukonthip Savatenalinton 2021-01-27 2021-01-27 733 19–41 19–41 10.5852/ejt.2021.733.1217 Nihonella gen. nov., a new troglophilic genus of dwarf spiders from Japan with a discussion on its phylogenetic position within the subfamily Erigoninae (Araneae, Linyphiidae) <p>A new monospecific genus belonging to the family Linyphiidae Blackwell, 1859, <em>Nihonella</em> gen.&nbsp;nov., is described using an integrative taxonomic approach based on the species <em>N.&nbsp;chika</em> gen.&nbsp;et&nbsp;sp.&nbsp;nov. The new genus is endemic to Western Honshu, Japan, and it shows distinctive genitalic and somatic characters of other genera of the subfamily Erigoninae Emerton, 1882. <em>Nihonella</em> gen.&nbsp;nov. is found only in the twilight and transition zones of caves in Okayama and Nara Prefectures. The phylogenetic position of <em>Nihonella</em> gen.&nbsp;nov. within the subfamily Erigoninae, and its relationship as a sister clade of the species of the group of <em>Savignia</em> Blackwell, 1833 (sensu Millidge 1977), is discussed on the basis of both, morphological and molecular evidence.</p> Francesco Ballarin Takeshi Yamasaki Copyright (c) 2021 Francesco Ballarin, Takeshi Yamasaki 2021-01-26 2021-01-26 733 1–18 1–18 10.5852/ejt.2021.733.1215 Revision of the morphology, phylogenetic relationships, behaviour and diversity of the Iberian and Italian ant-like Tachydromia Meigen, 1803 (Diptera: Hybotidae) <p>Phylogenetic inference, based on five molecular markers (COI, 28S, AATS, 12S, PGD), corroborates the synonymy of the flightless genera <em>Pieltainia</em> Arias, 1919 and <em>Ariasella</em> Gil, 1923 with <em>Tachydromia</em> Meigen, 1803. The secondary structure of the 28S rRNA gene is used for the first time in this family to align the multiple sequences. Molecular and morphological data are largely congruent for all known species of flightless <em>Tachydromia</em>. This paper treats ten western Mediterranean species (nine Iberian and one Italian) in detail, including the description of four new species: <em>T. ebejeri</em> Gonçalves, Grootaert &amp; Andrade sp. nov., <em>T. stenoptera</em> Gonçalves, Grootaert &amp; Andrade sp. nov., <em>T. cantabrica</em> Gonçalves, Grootaert &amp; Andrade sp. nov. and <em>T. nigrohirta</em> Gonçalves, Grootaert &amp; Andrade sp. nov. The male of <em>Tachydromia pieltaini</em> (Gil Collado, 1936) and the female of <em>Tachydromia apterygon</em> Plant &amp; Deeming, 2006 are described for the first time, while a lectotype is assigned to <em>Tachydromia pandellei</em> (Séguy, 1941). A key to all non-macropterous <em>Tachydromia</em> is supplied. Knowledge on the geographic distribution of most species is considerably enhanced. The mating behaviour of <em>Tachydromia semiaptera</em> (Gil Collado, 1923) and <em>Tachydromia iberica</em> (Arias, 1919) is documented for the first time, and we propose a change in the definition of terms apterous and micropterous to properly accommodate the diversity of wing states in this cluster of species.</p> Ana Rita Gonçalves Patrick Grootaert Rui Andrade Octávio S. Paulo Ximo Mengual Copyright (c) 2021 Ana Rita Gonçalves, Patrick Grootaert, Rui Andrade, Octávio S. Paulo, Ximo Mengual 2021-01-25 2021-01-25 733 1 56 10.5852/ejt.2021.732.1213 Mumetopia interfeles sp. nov., a new species of Anthomyzidae (Diptera) occurring en masse in an urban grassy habitat in Chile: its taxonomy, phylogeny and biology <p>A new species of Anthomyzidae (Diptera), viz., <em>Mumetopia interfeles</em> Roháček sp.&nbsp;nov. (both sexes), is described from Chile, based on a large series collected from a small grassy area (consisting of a non-native species, <em>Hordeum murinum</em> L.) among houses in the city of Valparaíso. Its systematic affiliation, phylogenetic relationships, and biology are discussed. Cases of mass occurrence of adult Anthomyzidae, particularly those in anthropogenic habitats, are reviewed and general features of them discussed. A phylogenetic hypothesis is proposed for the relationships of the new species and the <em>Chamaebosca</em> group of genera within Anthomyzidae (64 species in dataset), based on Bayesian inference and maximum likelihood analyses of seven DNA markers (12S, 16S, 28S, COI, COII, CytB, ITS2). Taxonomic limits of the genus <em>Mumetopia</em> Melander, 1913 are discussed. Based on a new molecular hypothesis and a previous cladistic analysis of morphological data (Roháček &amp; Barber 2009), a broadened, better-supported concept of <em>Mumetopia</em> (s.&nbsp;str.) is proposed to include <em>M.&nbsp;interfeles</em> sp.&nbsp;nov. (and its unnamed relatives) and externally aberrant, undescribed species endemic to the Juan Fernández Islands. Apart from Anthomyzidae (<em>M.&nbsp;interfeles</em> sp.&nbsp;nov. only), the community of Diptera associated with <em>H. murinum</em> was dominated by phytophagous Chloropidae: <em>Ceratobarys sacculicornis</em> (Enderlein, 1911) occurred in extremely high numbers similar to those of <em>M.&nbsp;interfeles</em> sp.&nbsp;nov., while the less abundant <em>Opetiophora</em> sp. represents the first record of the genus from South America.</p> Jindřich Roháček Andrea Tóthová Copyright (c) 2021 Jindřich Roháček, Andrea Tóthová 2021-01-21 2021-01-21 733 135 158 10.5852/ejt.2021.731.1211 Systematics of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) from the Andes of Northern Chile <p>The adult stage of <em>Helioandesia tarregai</em> gen.&nbsp;et&nbsp;sp.&nbsp;nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) is described and illustrated from the arid western slopes of the Andes of northern Chile. The larvae of <em>H.&nbsp;tarregai</em> gen.&nbsp;et&nbsp;sp.&nbsp;nov. feed as leaf skeletonizers on <em>Mirabilis acuta</em> (Reiche) Heimerl (Nyctaginaceae). The mostly gray forewing of <em>H.&nbsp;tarregai</em> gen.&nbsp;et&nbsp;sp.&nbsp;nov., ornamented with strongly bulging metallic spots, resembles that of the representatives of the mainly Nearctic <em>Lithariapteryx</em> Chambers, 1876. However, the latter lacks CuP in the forewing, has a single bristle in the female frenulum, and lacks a well-developed cornutus. <em>Helioandesia</em> gen.&nbsp;nov. clustered as sister to <em>Neoheliodines</em> Hsu, 2004 in a cladistic analysis, although no synapomorphies were found for this cluster, while <em>Lithariapteryx </em>was sister to<em> Helioandesia</em> gen.&nbsp;nov. + <em>Neoheliodines</em> based on two synapomorphies. The genetic distance between a DNA barcode sequence of <em>H.&nbsp;tarregai</em> gen.&nbsp;et&nbsp;sp.&nbsp;nov. and representatives of other genera of Heliodinidae Heinemann, 1877 was 9.0–12.5% (K2P), and a maximum likelihood analysis based on this molecular marker confirmed the placement of <em>H.&nbsp;tarregai</em> gen.&nbsp;et&nbsp;sp.&nbsp;nov. as a member of this micromoth family. This contribution represents the first confirmed record of Heliodinidae for Chile.</p> Héctor A. Vargas Copyright (c) 2021 Héctor A. Vargas 2021-01-20 2021-01-20 733 117–134 117–134 10.5852/ejt.2021.731.1209 A new spider genus (Araneae: Linyphiidae: Erigoninae) from a tropical montane cloud forest of Mexico <p>A new genus and species of spider (Araneae, Linyphiidae, Erigoninae) from a tropical montane cloud forest of Mexico is described from both male and female specimens, <em>Xim trenzado</em> gen.&nbsp;et sp. nov. A phylogenetic parsimony analysis situates <em>Xim</em> gen.&nbsp;nov. as a distinct genus among the distal Erigoninae. <em>Xim</em> gen.&nbsp;nov. is sister to a clade including <em>Ceratinopsis</em>, <em>Tutaibo</em> and <em>Sphecozone</em>, but differs from those genera by having a high cymbium, large paracymbium, short straight embolus, male cheliceral stridulatory striae widely and evenly spaced, both sexes with a post-ocular lobe, male with two series of prolateral macrosetae on femur I, and the female by having strongly oblong, u-shaped spermathecae.</p> Guillermo Ibarra-Núñez David Chamé-Vázquez Julieta Maya-Morales Copyright (c) 2021 Guillermo Ibarra-Núñez, David Chamé-Vázquez, Julieta Maya-Morales 2021-01-19 2021-01-19 733 97–116 97–116 10.5852/ejt.2021.731.1207 All-inclusive descriptions of new freshwater snail taxa of the hyperdiverse family Tateidae (Gastropoda, Caenogastropoda) from the South Island of New Zealand <p>Four new species and one new subspecies of tateid freshwater gastropods are described from the north of the South Island of New Zealand, <em>Catapyrgus jami</em> sp. nov., <em>Opacuincola lisannea</em> sp. nov., <em>O. gretathunbergae</em> sp. nov., <em>O. mete kahurangi</em> ssp. nov. and <em>Obtusopyrgus farri</em> sp. nov. The species are integratively defined based on a combination of shell morphological, anatomical and mitochondrial DNA data. Morphological and anatomical data were generated by morphometrics, scanning electron microscopy, as well as micro-computed tomography. The genetic data were basis of phylogenetic analyses and incorporated into the diagnoses. The new taxa occur in springs or spring-like habitats, i.e., shallow, slow-flowing sections of small streams except for <em>O. mete kahurangi</em> subsp. nov., which was collected from rough rocks in a river, where the snails sat in small depressions. None of the species exceeded 2.75 mm in length. <em>Opacuincola gretathunbergae</em> sp. nov. and <em>Obtusopyrgus farri</em> sp. nov. are pigmented and true crenobionts, while <em>C. jami</em> sp. nov. and the sympatric <em>Opacuincola lisannea</em> sp. nov. have eyes of reduced size and lack epidermal pigment, hence, probably dwell in the transitional zone of epigean and groundwaters.</p> Gerlien Verhaegen Martin Haase Copyright (c) 2021 Gerlien Verhaegen, Martin Haase 2021-01-18 2021-01-18 733 71 96 10.5852/ejt.2021.731.1205 A new Cyrtodactylus (Squamata: Gekkonidae) from Binh Thuan Province, southern Vietnam <p>We describe a new species of <em>Cyrtodactylus</em> on the basis of two specimens collected from Ta Kou Nature Reserve, Binh Thuan Province, southern Vietnam. <em>Cyrtodactylus chungi</em> sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: relatively small body size (SVL up to 68.5 mm); a continuous neckband; 5 or 6 irregular transverse dorsal bands; 11 or 12 bands on original tail; keeled tubercles present on dorsum, posterior limbs and tail; 17 or 18 irregular dorsal tubercle rows; 30 or 31 ventral scale rows; ventrolateral skin folds indistinct; an angular series of seven precloacal pores in male and six pitted, enlarged precloacal scales in female, each series separated by a diastema of undifferentiated scales from 4–6 enlarged, poreless femoral scales; median subcaudals slightly enlarged; 17–20 subdigital lamellae under the fourth toe. Based on molecular analyses of the fragment of mitochondrial gene cytochrome c oxidase subunit I (COI), the new species is recovered as the sister taxon to <em>Cyrtodactylus cattienensis</em> s. str. with a genetic divergence of more than 9%. In phylogenetic analyses, the new species is recovered as a member of the <em>Cyrtodactylus irregularis</em> species group.</p> Sabrina Ostrowski Minh Duc Le Hanh Thi Ngo Cuong The Pham Trung My Phung Truong Quang Nguyen Thomas Ziegler Copyright (c) 2021 Sabrina Ostrowski, Minh Duc Le, Hanh Thi Ngo, Cuong The Pham, Trung My Phung, Truong Quang Nguyen, Thomas Ziegler 2021-01-13 2021-01-13 733 47 70 10.5852/ejt.2021.731.1203 Asian cave-adapted diplurans, with the description of two new genera and four new species (Arthropoda, Hexapoda, Entognatha) <p>Twenty-nine specimens of Diplura collected from eight caves in China and Myanmar contain two new genera, <em>Hubeicampa</em> Sendra &amp; Lips gen. nov. and <em>Mueggejapyx</em> Sendra &amp; Komerički gen. nov., as well as four new species, <em>Anisuracampa ywangana</em> Sendra &amp; Komerički sp. nov., <em>Hubeicampa melissa</em> Sendra &amp; Lips gen. et sp. nov., <em>Pacificampa wudonghuii</em> Sendra sp. nov. and <em>Mueggejapyx brehieri</em> Sendra &amp; Komerički gen. et sp. nov. These cave-adapted taxa showcase an impressive diversity in morphological adaptation (troglomorphy) to cave ecosystems. Their sensorial equipment, setae and receptors in the cupuliform organ have unique forms (<em>H. melissa</em> gen. et sp. nov.), as well as the pretarsus sticky surface (<em>A. ywangana</em> sp. nov. and <em>H. melissa</em> gen. et sp. nov.). Recent contributions on Asian diplurans, together with the taxonomic novelties shown in the present study, highlight the biogeographical importance of the Asian biodiversity. Asia is revealed as a continent with vast karst regions still waiting to be explored and new dipluran species waiting to be discovered.</p> Alberto Sendra Ana Komerički Josiane Lips Yunxia Luan Jesús Selfa Alberto Jiménez-Valverde Copyright (c) 2021 Alberto Sendra, Ana Komerički, Josiane Lips, Yunxia Luan, Jesús Selfa, Alberto Jiménez-Valverde 2021-01-05 2021-01-05 733 1 46 10.5852/ejt.2021.731.1199