European Journal of Taxonomy <p>The <em>European Journal of Taxonomy</em> is a peer-reviewed international journal in descriptive taxonomy, covering the eukaryotic world. Its content is fully electronic and <a href="">Open Access</a>. It is published and funded by a <a href="">consortium</a> of European natural history institutions. Neither authors nor readers have to pay fees. All articles published in <em>EJT</em> are compliant with the different nomenclatural codes. <em>EJT</em> is an archived and indexed journal that welcomes scientific contributions from all over the world, both in content and authorship. If you have any questions about <em>EJT</em>, please <a href="">contact us</a></p> en-US <h3>Creative Commons Copyright Notices</h3> <div class="page"> <p>Authors who publish with this journal agree to the following terms:</p> <ol> <li class="show">Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a&nbsp;<a href="">Creative Commons Attribution License</a> (CC BY 4.0) that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.</li> <li class="show">Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.</li> <li class="show">Authors are NOT ALLOWED TO post their work online (e.g., in institutional repositories or on their website) prior to and during the submission process, as it can lead to taxonomic issues.</li> </ol> </div> (Laurence Benichou) (Kristiaan Hoedemakers) Tue, 21 Sep 2021 01:51:12 -0700 OJS 60 Morphology and Taxonomy of Isopoda Anthuroidea (Crustacea) from Sulawesi with description of six new species <p>This is a first inventory of Isopoda Anthuroidea from near-shore marine sites in Sulawesi. The material was collected from reefs, sea-grass beds and mangroves of the Minahasa Peninsula in North Sulawesi. Specimens were washed out from benthos samples collected in shallow water. The new species <em>Hyssura reptans </em>sp. nov.<em>, Kupellonura indonesica </em>sp. nov.<em>, Kupellonura macaroni </em>sp. nov.<em>, Pendanthura bangkaensis </em>sp. nov.<em>, Expanathura marcoi </em>sp. nov.<em>, </em>and<em> Colanthura</em> <em>gerungi </em>sp. nov. are described. We discovered that in <em>Colanthura </em>the mandible is fused to the cephalothorax, a unique feature among isopods. <em>Expanathura collaris </em>(Kensley, 1979) is described from Bangka Island and several differences between other Indo-Pacific populations are discussed.</p> Manikmayang Annisaqois, J. Wolfgang Wägele Copyright (c) 2021 Manikmayang Annisaqois, J. Wolfgang Wägele Tue, 21 Sep 2021 01:46:19 -0700 Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902) <p>Five new species of the terrestrial snail genus <em>Landouria</em> Godwin-Austen, 1918 (Camaenidae) are described from northeastern Thailand, based on shell features, radular morphology, genital anatomy, and DNA sequence data: <em>Landouria circinata</em> sp. nov., <em>L. tuberculata</em> sp. nov., <em>L. trochomorphoides</em> sp. nov., <em>L. chloritoides</em> sp. nov., and <em>L. elegans</em> sp. nov. These species are phylogenetically well separated from each other by mtDNA phy-logeny and COI sequence divergences of 0.073–0.156. The record of <em>Thaitropis</em> Schileyko, 2004 (currently synonymized with <em>Landouria</em>) in Thailand is re-interpreted as referring to <em>L. diplogramma</em> (Möllendorff, 1902) comb. nov.</p> Benchawan Nahok, Sakboworn Tumpeesuwan, Chanidaporn Tumpeesuwan Copyright (c) 2021 Benchawan Nahok, Sakboworn Tumpeesuwan, Chanidaporn Tumpeesuwan Fri, 17 Sep 2021 01:44:56 -0700 A review of Paratranes Zimmerman, 1994, Xanthorrhoea-associated weevils of the Tranes group (Coleoptera, Curculionidae, Molytinae), with description of a new species <p>Grasstrees (<em>Xanthorrhoea</em>) are a group of monocotyledonous plants endemic to Australia, with a characteristic crown of long and narrow leaves. In the present study, the grasstree-associated weevils of the genus <em>Paratranes</em> Zimmerman, 1994 are reviewed and two species are recognised: <em>P.&nbsp;monopticus</em> (Pascoe, 1870) and <em>P.&nbsp;zimmermani</em> sp.&nbsp;nov. A lectotype for <em>Tranes xanthorrhoeae</em> Lea, 1898, which was recently synonymised with <em>P.&nbsp;monopticus</em>, is herein designated, and the validity of this synonymy is confirmed. Descriptions and diagnoses of the species are supplemented with photographs of their habitus and salient structures as well as a map of their distributions.</p> Yun Hsiao, Rolf G. Oberprieler Copyright (c) 2021 Yun Hsiao, Rolf G. Oberprieler Thu, 16 Sep 2021 02:46:52 -0700 A contribution to the genus Afronurus Lestage, 1924 in China (Ephemeroptera: Heptageniidae, Ecdyonurinae) <p>The genus <em>Afronurus</em> has several very common mayfly species in China and they are widely distributed in this country. Some of them are quite similar to each other in both imaginal and nymphal stages. However, these species have not been systematically compared and reviewed so far. In this study, six species are recognized. All nymphs of them share the following characters: gills V–VI with additional arrow-like accessory lobes, branched dentisetae, two rows of bristles and setae on hindtibiae and spotted abdominal terga. The males have divergent penes and clearly expressed titillators. The nymphs of the new species <em>A.&nbsp;drepanophyllus </em>sp. nov. have sickle-like gills I, spotted and striped body color, and males have unique genitalia. The nymphal stages of <em>A.&nbsp;furcatus</em> and <em>A.&nbsp;hunanensis</em>, which are associated and described for the first time, have similar body color to <em>A.&nbsp;drepanophyllus </em>sp. nov., but their pale dots on the head capsules and the shape of the hypopharynx are different. Keys to males and nymphs of the six species are provided.</p> Wei Zhang, Zhi-Ming Lei, Wen-Juan Li, Chang-Fa Zhou Copyright (c) 2021 Wei Zhang, Zhi-Ming Lei, Wen-Juan Li, Chang-Fa Zhou Wed, 15 Sep 2021 03:29:03 -0700 Two new species of the planthopper genus Eponisiella Emeljanov from China (Hemiptera, Fulgoromorpha, Meenoplidae) <p>Two new species of the genus <em>Eponisiella</em> Emeljanov, 1984 are described and illustrated from China (Guizhou and Shandong Provinces). These are <em>E.&nbsp;dafangensis</em> sp.&nbsp;nov. and <em>E.&nbsp;shandongensis</em> sp.&nbsp;nov., giving the genus eight species in total. A key to the species of <em>Eponisiella</em> is provided as well as a map of their geographic distribution, which is briefly discussed.</p> Sha-sha Lv, Lin Yang, Xiang-Sheng Chen Copyright (c) 2021 Sha-sha Lv, Lin Yang, Xiang-Sheng Chen Tue, 14 Sep 2021 00:59:37 -0700 Systematic revision of the limestone karst-restricted land snail genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with description of a new species <p>Thai limestone karsts are known to contain a rich biodiversity of animals, especially terrestrial snails, but still require further intensive exploration to evaluate their biodiversity. To date, only a few studies on the limestone karst-inhabiting land snail genera have been published. The present work focuses on the species diversity and phylogenetic relationships of the limestone karst-restricted land snail genus <em>Aenigmatoconcha</em> from Thailand, based on comparative morphology and molecular evidence. The results yielded three known species (<em>A. clivicola</em> Tumpeesuwan &amp; Tumpeesuwan, 2017, <em>A. sumonthai</em> Tumpeesuwan &amp; Tumpeesuwan, 2018, and <em>A. mitis</em> (Pfeiffer, 1863) comb. nov.), plus a new species (<em>A. eunetis</em> Pholyotha &amp; Panha sp. nov). The phylogenetic analyses of partial fragments of the mitochondrial cytochrome oxidase c subunit I (COI) gene confirmed the monophyly of all recognized species and congruence with the traditional morphology-based species designations. Average uncorrected p-distances of COI sequences between species were 9.7–12.0% and within species were 0.2–4.2%. This study also provides the re-description of penial sculpture, penial sheath, flagellum, penial caecum, and mantle lobe morphology that were neglected from the type species description. The present discovery of a new species increases the known diversity of Thai land snails and will support the conservation planning to protect karst biodiversity.</p> Arthit Pholyotha, Chirasak Sutcharit, Piyoros Tongkerd, Somsak Panha Copyright (c) 2021 Arthit PHOLYOTHA, Chirasak SUTCHARIT, Piyoros TONGKERD, Somsak PANHA Fri, 10 Sep 2021 07:37:21 -0700 Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819 <p>The scarab beetles of the subfamily Orphninae (Coleoptera: Scarabaeidae) from Sri Lanka are reviewed. Four species of the genus <em>Orphnus</em> Macleay, 1819, are recorded from the island: <em>O. bicolor</em> (Fabricius, 1801), <em>O. parvus</em> (Wiedemann, 1823), <em>O. mysoriensis</em> Westwood, 1845, and <em>O. medvedevi</em> sp. nov. Lectotypes are designated for the three former names. Synonymy of <em>O. detegens</em> Walker, 1859, and <em>O. scitissimus</em> Walker, 1859, is discussed. Keys, illustrations of habitus and male genitalia, and distributional record maps are given for all species.</p> Andrey V. Frolov, Lilia A. Akhmetova Copyright (c) 2021 Andrey V. Frolov, Lilia A. Akhmetova Mon, 06 Sep 2021 01:58:03 -0700 Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae) <p>Three new species of <em>Patrus</em> Aubé, 1838 are described from China: <em>Patrus</em> <em>hainanensis</em> sp.&nbsp;nov. from Hainan; <em>Patrus jiangxiensis </em>sp.&nbsp;nov. from Jiangxi; <em>Patrus shangchuanensis</em> sp.&nbsp;nov. from Guangdong. Eight species / subspecies of Gyrinidae are recorded from China for the first time: <em>Metagyrinus vitalisi </em>(Peschet, 1923), <em>Orectochilus</em> <em>argenteolimbatus</em> Peschet, 1923, <em>Orectochilus murinus</em> Régimbart, 1892, <em>Patrus haemorrhous </em>(Régimbart, 1892), <em>Patrus marginepennis angustilimbus </em>(Ochs, 1925) from Yunnan; <em>Patrus coomani </em>(Peschet, 1925) from Guangdong; <em>Patrus procerus</em> (Régimbart, 1884) from Guangxi;<em> Patrus annandalei</em> (Ochs, 1925) from Hainan. Additional faunistic data of Gyrinidae from China are provided. A key to Chinese species of <em>Patrus</em> Aubé based on examined specimens from China is given.</p> Zulong Liang, Robert B. Angus, Fenglong Jia Copyright (c) 2021 Zulong Liang, Robert B. Angus, Fenglong Jia Wed, 01 Sep 2021 01:32:37 -0700 Interactive identification key to all brittle star families (Echinodermata; Ophiuroidea) leads to revised morphological descriptions <p>Ophiuroidea is the largest class among extant echinoderms, with over 2000 described species assigned to 33 families. Here, the first identification key to the recently revised classification was developed, and revised morphological descriptions were derived from it, expanding the previous short diagnoses. The key was built by analyzing internal and external skeletal characters of predominantly the type species of each family, including at least two mutually exclusive attributes per family. Various numeric and multistate characters were used to create a traditional as well as an interactive key using the DELTA and Xper software programs­. Illustrations (SEM and digital photos) are included in the key to facilitate the assessment of character states by users. Not only is it the first identification key to the families, according to the recently proposed new classification and the examined species, but this interactive key also assists users in understanding the family level taxonomy of brittle stars. The interactive key allows new characters and states to be added, when more species will have been analyzed, without the need to reconfigure the complete key (as may be necessary with conventional keys).</p> Mona Goharimanesh, Sabine Stöhr, Omid Mirshamsi, Fereshteh Ghassemzadeh, Dominique Adriaens Copyright (c) 2021 Mona Goharimanesh, Sabine Stöhr, Omid Mirshamsi, Fereshteh Ghassemzadeh, Dominique Adriaens Tue, 31 Aug 2021 00:00:00 -0700 Review of the braconid parasitoid subfamily Doryctinae (Hymenoptera, Braconidae) from the United Arab Emirates and Yemen <p>Yemen is provided for the first time. The following genera are recorded in the southern Arabian Peninsula for the first time: tribe Doryctini – <em>Hemispathius</em> Belokobylskij &amp; Quicke, 2000 and <em>Doryctes</em> Haliday, 1836; tribe Spathiini – <em>Parana</em> Nixon, 1941 and <em>Spathius</em> Nees, 1819; tribe Hecabolini – <em>Hemidoryctes</em> Belokobylskij, 1992 and <em>Parallorhogas</em> Marsh, 1993; tribe Heterospilini –<em> Heterospilus</em> Haliday 1936; tribe Rhaconotini – <em>Platyspathius</em> Viereck, 1911 and <em>Rhaconotinus</em> Hedqvist, 1965. Sixteen species and one subspecies are described as new for science: <em>Dendrosotinus</em> (<em>Gildoria</em>) <em>maculipennis</em> Belokobylskij sp.&nbsp;nov., <em>D.</em>&nbsp;(<em>G.</em>)&nbsp;<em>subelongatus</em> Belokobylskij sp.&nbsp;nov., <em>Doryctes</em> (<em>Neodoryctes</em>) <em>arrujumi</em> Belokobylskij sp.&nbsp;nov., <em>Parana arabica</em> Belokobylskij sp.&nbsp;nov., <em>Spathius alkadanus</em> Belokobylskij sp.&nbsp;nov., <em>S.</em>&nbsp;<em>austroarabicus</em> Belokobylskij sp.&nbsp;nov., <em>S. lahji</em> Belokobylskij sp.&nbsp;nov., <em>S. subafricanus</em> Belokobylskij sp.&nbsp;nov., <em>Hecabalodes</em> <em>maculatus</em> Belokobylskij sp.&nbsp;nov., <em>Platyspathius (Platyspathius) longicaudis</em> Belokobylskij sp.&nbsp;nov., <em>P. </em>(<em>P.</em>)<em> brevis</em> Belokobylskij sp.&nbsp;nov., <em>Rhaconotinus albosetosus</em> Belokobylskij sp.&nbsp;nov., <em>Rhaconotus brevicellularis</em> Belokobylskij sp.&nbsp;nov., <em>Rh.&nbsp;magniareolus</em> Belokobylskij sp.&nbsp;nov., <em>Rh.&nbsp;microexcavatus </em>Belokobylskij sp.&nbsp;nov., <em>Rh.&nbsp;vanharteni</em> Belokobylskij sp.&nbsp;nov. and <em>Hemidoryctes</em> <em>carbonarius postfurcalis</em> Belokobylskij subsp.&nbsp;nov. Two new generic combinations are proposed: <em>Hemispathius pilosus </em>(Granger, 1949) comb. nov. (transferred from <em>Doryctes</em>) and <em>Parallorhogas testaceus</em> (Szépligeti, 1914) comb. nov. (transferred from <em>Opius</em>). <em>Rhaconotus decaryi</em> Granger, 1949 is here synonymised with <em>Rh.&nbsp;menippus </em>Nixon, 1939 (syn.&nbsp;nov.). A lectotype for <em>Doryctes</em> <em>pilosus </em>Granger, 1949 is designated. The following species are recorded for the UAE and/or Yemen for the first time: <em>Dendrosotinus</em> <em>ferrugineus</em> (Marshall, 1888), <em>Hemispathius pilosus </em>(Granger, 1949), <em>Mimodoryctes</em> <em>proprius</em> Belokobylskij, 2001, <em>M. arabicus</em> Edmardash, Gadallah &amp; Soliman, 2020, <em>Spathius</em> <em>nixoni</em> Belokobylskij &amp; Maetô, 2009, <em>Hecabalodes anthaxiae</em> Wilkinson, 1929, <em>H. radialis</em> Tobias, 1962, <em>H.&nbsp;xylophagi</em> Fischer, 1962, <em>Parallorhogas testaceus</em> (Szépligeti, 1914), <em>Heterospilus</em> (<em>Eoheterospilus</em>) <em>rubrocinctus</em> (Ashmead, 1905), <em>Rhaconotinus menippus </em>(Nixon, 1939), <em>Rhaconotus arabicus</em> Belokobylskij, 2001, <em>Rh.&nbsp;manolus</em> Nixon, 1941, <em>Rh.&nbsp;scirpophagae</em> Wilkinson, 1927 and <em>Rh.&nbsp;sudanensis </em>Wilkinson, 1927.</p> Sergey A. Belokobylskij, Cornelis van Achterberg Copyright (c) 2021 Sergey A. Belokobylskij, Cornelis van Achterberg Mon, 30 Aug 2021 01:52:31 -0700