Diversity and taxonomy of the Late Triassic sauropodomorphs (Saurischia, Sauropodomorpha) stored in the Palaeontological Collection of Tübingen, Germany, historically referred to Plateosaurus
Abstract
The genus Plateosaurus is one of the most challenging aspects of early-diverging sauropodomorph taxonomy, with a total of 29 names and a century of revisions. As a result, nomina dubia have been treated as synonyms of Plateosaurus trossingensis, adding to the confusion about the extent of morphological variability. Here, we provide a thorough revision of the taxonomy proposed by von Huene as well as the subsequent taxonomic interpretations of Plateosaurus through a complete inventory of the specimens stored in the Palaeontological Collection of Tübingen. We reassess the status of the specimens in the collection and propose that the sauropodomorph-bearing layers are not necessarily monospecific. Most of the original fieldwork documentation has been lost, but we use the taxonomy established by von Huene as a historical reference point to reconstruct what was known at the time. This revised taxonomy of Plateosaurus narrows the genus to three species: Pl. trossingensis, Pl. longiceps, and Pl. gracilis (as a metataxon), and restricts the genera Gresslyosaurus and Pachysaurus to large and robust individuals, as pragmatic decisions aimed to test their affinities to other Late Triassic sauropodomorphs. Future studies should consider not only morphological variability, but also stratigraphy, palaeogeography, and environmental data when delineating species within and outside the Plateosaurus plexus.
References
Agassiz L. 1846. Nomenclatoris zoologici index universalis: continens momina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Sumptibus Jent et Gassman, Soloduri [Solothurn].
Allain R. & Pereda Suberbiola X. 2003. Dinosaurs of France. Comptes Rendus Palevol 2 (1): 27–44. https://doi.org/10.1016/s1631-0683(03)00002-2
Allmon W.D. 2013. Species, speciation and palaeontology up to the Modern Synthesis: persistent themes and unanswered questions. Palaeontology 56 (6): 1199–1223. https://doi.org/10.1111/pala.12054
Anonymous 1847. Zweite Generalversammlung am 1. Mai 1846 zu Tübingen. Jahreshefte des Vereins für vaterländische Naturkunde in Württemberg: 129–184.
Barth G., Franz M., Heunisch C., Kustatscher E., Thies D., Vespermann J. & Wolfgramm M. 2014. Late Triassic (Norian-Rhaetian) brackish to freshwater habitats at a fluvial-dominated delta plain (Seinstedt, Lower Saxony, Germany). Palaeobiodiversity and Palaeoenvironments 94: 495–528. https://doi.org/10.1007/s12549-014-0168-6
Beccari V., Mateus O., Wings O., Milàn J. & Clemmensen L.B. 2021. Issi saaneq gen. et sp. nov.—A new sauropodomorph dinosaur from the Late Triassic (Norian) of Jameson Land, Central East Greenland. Diversity 13 (11): 561. https://doi.org/10.3390/d13110561
Benton M.J. 1986. The Late Triassic reptile Teratosaurus – A rauisuchian, not a dinosaur. Palaeontology 29 (2): 293–301.
Benton M.J. 1993. Late Triassic terrestrial vertebrate extinctions: stratigraphic aspects and the record of the Germanic Basin. In: Mazin J.M. &Pinna G. (eds) Evolution, Ecology and Biogeography of the Triassic Reptiles: 35–38. Società Italiana di Scienze Naturali, Museo Civico di Storia Naturale di Milano, Milan.
Benton M.J., Bernardi M. & Kinsella C. 2018. The Carnian Pluvial Episode and the origin of dinosaurs. Journal of the Geological Society 175 (6): 1019–1026. https://doi.org/10.1144/jgs2018-049
Bernardi M., Gianolla P., Petti F.M., Mietto P. & Benton M.J. 2018. Dinosaur diversification linked with the Carnian Pluvial Episode. Nature Communications 9 (1): 1499. https://doi.org/10.1038/s41467-018-03996-1
Bloos G., Dietl G. & Schweigert G. 2006. Der Jura Süddeutschlands in der Stratigraphischen Tabelle von Deutschland 2002. Newsletters on Stratigraphy 41: 263–277. https://doi.org/10.1127/0078-0421/2005/0041-0263
Böhmer C., Rauhut O.W.M. & Wörheide G. 2015. Correlation between Hox code and vertebral morphology in archosaurs. Proceedings of the Royal Society B: Biological Sciences 282 (1810): 20150077. https://doi.org/10.1098/rspb.2015.0077
Brusatte S.L., Butler R.J., Sulej T. & Niedźwiedzki G. 2009. The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland. Acta Palaeontologica Polonica 54 (2): 221–230. https://doi.org/10.4202/app.2008.0065
Buckland W. 1824. Notice on the Megalosaurus or great Fossil Lizard of Stonesfield. Transactions of the Geological Society of London 2 (1): 390–396. https://doi.org/10.1144/transgslb.1.2.390
Charig A.J., Attridge J. & Crompton A.W. 1965. On the origin of the sauropods and the classification of the Saurischia. Proceedings of the Linnean Society of London 176 (2): 197–221. https://doi.org/10.1111/j.1095-8312.1965.tb00944.x
Clemens W.A. & Martin T. 2014. Review of the non-tritylodontid synapsids from bone beds in the Rhaetic Sandstone, southern Germany. Paläontologische Zeitschrift 88 (4): 461–479. https://doi.org/10.1007/s12542-013-0201-5
Clemmensen L.B., Milàn J., Adolfssen J.S., Estrup E.J., Frobøse N., Klein N., Mateus O. & Wings O. 2016. The vertebrate-bearing Late Triassic Fleming Fjord Formation of central East Greenland revisited: Stratigraphy, palaeoclimate and new palaeontological data. Geological Society Special Publication 434 (1): 31–47. https://doi.org/10.1144/SP434.3
Cuny G. & Mazin J.M. 1993. The Late Triassic (Upper Keuper) dinosaur locality of Lons-le-Saunier (Jura, France): first results of excavations. Revue de Paléobiologie 7 (vol. spéc.): 45–53.
Daudin F.M. 1802. Histoire naturelle, génerale et particulière des Reptiles; Ouvrage faisant suite à l'Histoire naturelle, générale et particulière, composée par Leclerc de Buffon, et redigée par C.S. Sonnini, membre de plusieurs Sociétés savantes. Vol. 3. F. Dufart, Paris. https://doi.org/10.5962/bhl.title.60678
Etzold A. & Schweizer V. 2005. Der Keuper in Baden-Württemberg. In: Deutsche Stratigraphische Kommission (ed.) Stratigraphie von Deutschland IV, Keuper. Courier Forschungsinstitut Senckenberg 253: 214–258. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart.
Fitzinger L.J. 1843. Systema reptilium Fasciculus Primus: Amblyglossae. Braumüller et Seidel, Vienna. https://doi.org/10.5962/bhl.title.4694
Fraas E. 1896. Die Schwäbischen Trias-Saurier nach dem Material der Kgl. Naturalien-Sammlung in Stuttgart zusammengestellt. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart.
Franz M., Bachmann G.H., Barnasch J., Heunisch C. & Röhling H.-G. 2018. The Keuper Group in the Stratigraphic Table of Germany 2016 – continuous sedimentation in the North German Basin (variant B). Zeitschrift der Deutschen Gesellschaft für Geowissenschaften 169: 203–224. https://doi.org/10.1127/zdgg/2018/0114
Galton P.M. 1971. The prosauropod dinosaur Ammosaurus, the crocodile Protosuchus, and their bearing on the age of the Navajo Sandstone of northeastern Arizona. Journal of Paleontology 45 (5): 781–795.
Galton P.M. 1973. On the anatomy and relationships of Efraasia diagnostica (Huene) n. gen., a prosauropod dinosaur (Reptilia: Saurischia) from the Upper Triassic of Germany. Paläontologische Zeitschrift 47 (3/4): 229–255. https://doi.org/10.1007/bf02985709
Galton P.M. 1984a. An early prosauropod dinosaur from the Upper Triassic of Nordwürttemberg, West Germany. Stuttgarter Beiträge zur Naturkunde 106 (90): 1–25.
Galton P.M. 1984b. Cranial anatomy of the prosauropod dinosaur Plateosaurus from the Knollenmergel (Middle Keuper, Upper Triassic) of Germany. I. Two complete skulls from Trossingen/Wurtt, with comments on the diet. Geologica et Palaeontologica 18: 139–171.
Galton P.M. 1985a. Cranial anatomy of the prosauropod dinosaur Plateosaurus from the Knollenmergel (Middle Keuper, Upper Triassic) of Germany. II. All the material and details of soft-part anatomy. Geologica et Palaeontologica 19: 119–159.
Galton P.M. 1985b. Cranial anatomy of the prosauropod dinosaur Sellosaurus gracilis from the Middle Stubensandstein (Upper Triassic) of Nordwürttemberg, West Germany. Stuttgarter Beiträge zur Naturkunde 118: 1–39.
Galton P.M. 1985c. The poposaurid thecodontian Teratosaurus suevicus v. Meyer, plus referred specimens mostly based on prosauropod dinosaurs, from the Middle Stubensandstein (Upper Triassic) of Nordwürttemberg. Stuttgarter Beiträge zur Naturkunde 110: 1–29.
Galton P.M. 1986. Prosauropod dinosaur Plateosaurus (= Gresslyosaurus) (Saurischia: Sauropodornorpha) from the Upper Triassic of Switzerland. Geologica et Palaeontologica 20: 167–183.
Galton P.M. 1990. Basal Sauropodomorpha – Prosauropoda. In: Weishampel D.B., Dodson P. & Osmolska H. (eds) The Dinosauria. 1st Edition: 320–344. University of California Press, Berkeley and Los Angeles.
Galton P.M. 1998. The prosauropod dinosaur Plateosaurus (Dimodosaurus) poligniensis (Pidancet & Chopard, 1862) (Upper Triassic, Poligny, France). Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 207 (2): 255–288. https://doi.org/10.1127/njgpa/207/1998/255
Galton P.M. 1999. Sex, sacra and Sellosaurus gracilis (Saurischia, Sauropodomorpha, Upper Traissic, Germany) – or why the character ‘two sacral vertebrae’ is plesiomorphic for Dinosauria. Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 213 (1): 19–55. https://doi.org/10.1127/njgpa/213/1999/19
Galton P.M. 2000. The prosauropod dinosaur Plateosaurus Meyer, 1837 (Saurischia: Sauropodomorpha). I. The syntypes of P. engelhardti Meyer, 1837 (Upper Triassic, Germany), with notes on other European prosauropods with ‘distally straight’ femora. Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 216 (2): 233–275. https://doi.org/10.1127/njgpa/216/2000/233
Galton P.M. 2001a. The prosauropod dinosaur Plateosaurus Meyer, 1837 (Saurischia: Sauropodomorpha; Upper Triassic). II. Notes on the referred species. Revue de Paléobiologie 20 (2): 435–502.
Galton P.M. 2001b. Prosauropod dinosaurs from the Upper Triassic of Germany. In: Colectivo Arqueológico-Paleontológico de Salas, C.A.S (ed.) Actas de las I Jornadas Internacionales sobre Paleontología de Dinosaurios y su entorno. Salas de los Infantes (Burgos, España), septiembre de 1999: 25–92. Colectivo Arqueológico y Paleontológico de Salas, Burgos, Spain.
Galton P.M. 2001c. Prosauropod dinosaur Sellosaurus gracilis (Upper Triassic, Germany): Third sacral vertebra as either a dorsosacral or a caudosacral. Neues Jahrbuch für Geologie und Paläontologie – Monatshefte 2001 (11): 688–704. https://doi.org/10.1127/njgpm/2001/2001/688
Galton P.M. 2005. Bones of large dinosaurs (Prosauropoda and Stegosauria) from the Rhaetic Bone Bed (Upper Triassic) of Aust Cliff, southwest England. Revue de Paléobiologie 24 (1): 51–74.
Galton P.M. 2012. Case 3560 Plateosaurus engelhardti Meyer, 1837 (Dinosauria, Sauropodomorpha): proposed replacement of unidentifiable name-bearing type by a neotype. The Bulletin of Zoological Nomenclature 69 (3): 203–212. https://doi.org/10.21805/bzn.v69i3.a15
Galton P.M. 2013. Comment on Plateosaurus Meyer, 1837 (Dinosauria, Sauropodomorpha): proposed replacement of unidentifiable name-bearing type by a neotype (Case 3560; see BZN 69: 203–212, 295–296; 70: 120–121). Bulletin of Zoological Nomenclature 70 (3): 205–206.
Galton P.M. & Bakker R.T. 1985. The cranial anatomy of the prosauropod dinosaur ‘Efraasia diagnostica’, a juvenile individual of Sellosaurus gracilis from the Upper Triassic of Nordwürttemberg, West Germany. Stuttgarter Beiträge zur Naturkunde 117: 1–15.
Galton P.M. & Upchurch P. 2004. Prosauropoda. In: Weishampel D.B., Dodson P. & Osmólska H. (eds) The Dinosauria (2nd Edition): 232–258. University of California Press, Berkeley.
Galton P.M., Yates A.M. & Kermack D. 2007. Pantydraco n. gen. for Thecodontosaurus caducus Yates, 2003, a basal sauropodomorph dinosaur from the Upper Triassic or Lower Jurassic of South Wales, UK. Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 243 (1): 119–125. https://doi.org/10.1127/0077-7749/2007/0243-0119
Geyer O.F. & Gwinner M.P. 2011. Geologie von Baden-Württemberg. Schweizerbart’sche Verlagsbuch-handlung, Stuttgart.
Glut D.F. 1997. Dinosaurs. The Encyclopedia. First Edition. McFarland & Co., Jefferson, N.C.
Hofmann R. & Sander P.M. 2014. The first juvenile specimens of Plateosaurus engelhardti from Frick, Switzerland: isolated neural arches and their implications for developmental plasticity in a basal sauropodomorph. PeerJ 2: e458. https://doi.org/10.7717/peerj.458
Holz M. & Schultz C.L. 1998. Taphonomy of the south Brazilian Triassic herpetofauna: fossilization mode and implications for morphological studies. Lethaia 31 (4): 335–345. https://doi.org/10.1111/j.1502-3931.1998.tb00523.x
Hungerbühler A. 1998. Taphonomy of the prosauropod dinosaur Sellosaurus, and its implications for carnivore faunas and feeding habits in the Late Triassic. Palaeogeography, Palaeoclimatology, Palaeoecology 143 (1–3): 1–29. https://doi.org/10.1016/S0031-0182(98)00074-1
Hungerbühler A. 2001. The status and phylogenetic relationships of “Zanclodon” arenaceus: the earliest known phytosaur? Paläontologische Zeitschrift 75: 97–112. https://doi.org/10.1007/BF03022600
ICZN. 2019. Opinion 2435 (Case 3560) – Plateosaurus Meyer, 1837 (Dinosauria, Sauropodomorpha): new type species designated. The Bulletin of Zoological Nomenclature 76 (1): 144–145. https://doi.org/10.21805/bzn.v76.a042
Jenkins F.A., Shubin N.H., Amaral W.W., Gatesby S.M., Schaff C.R., Clemmensen L.B., Downs W.R., Davidson A.R., Bonde N. & Osbaeck F. 1994. Late Triassic Continental Vertebrates and Depositional Environments of the Fleming Fjord Formation, Jameson Land, East Greenland. Kommissionen for Videnskabelige Undersøgelser i Grønland, Copenhagen.
Jordan P., Pietsch J.S., Bläsi H., Furrer H., Kündig N., Looser N., Wetzel A. & Deplazes G. 2016. The middle to late Triassic Bänkerjoch and Klettgau formations of northern Switzerland. Swiss Journal of Geosciences 109 (2): 257–284. https://doi.org/10.1007/s00015-016-0218-3
Klein N. 2004. Bone Histology and Growth of the Prosauropod Dinosaur Plateosaurus engelhardti MEYER, 1837 from the Norian Bonebeds of Trossingen (Germany) and Frick (Switzerland). PhD thesis, Rheinische Friedrich-Wilhelms-Universität Bonn, Germany.
Kozur H.W. & Weems R.E. 2010. The biostratigraphic importance of conchostracans in the continental Triassic of the northern hemisphere. Geological Society, London, Special Publications 334 (1): 315–417. https://doi.org/10.1144/SP334.13
König C.D.E. 1818. Synopsis of the Contents of the British Museum. 13th Edition. British Museum, London.
Kuhn O. 1959. Ein neuer Microsaurier aus dem deutschen Rotliegenden. Neues Jahrbuch fur Geologie und Paläontologie Monatshefte 9: 424–426.
Lallensack J., Teschner E., Pabst B. & Sander P.M. 2021. New skulls of the basal sauropodomorph Plateosaurus trossingensis from Frick, Switzerland: is there more than one species? Acta Palaeontologica Polonica 66 (1): 1–28. https://doi.org/10.4202/app.00804.2020
Lucas S.G., Tanner L.H., Kozur H.W., Weems R.E. & Heckert A.B. 2012. The Late Triassic timescale: age and correlation of the Carnian–Norian boundary. Earth-Science Reviews 114 (1): 1–18. https://doi.org/10.1016/j.earscirev.2012.04.002
Maisch M.W. 2020. Friedrich von Huene (1875–1969) und seine Beiträge zur Alphataxonomie der Dinosaurier (Reptilia: Saurischia, Ornithischia) – eine kritische Würdigung. Jahreshefte der Gesellschaft für Naturkunde in Württemberg 176: 117–219. https://doi.org/10.26251/jhgfn.176.2020.117-219
Mallison H. 2010. The digital Plateosaurus I: Body mass, mass distribution and posture assessed using cad and cae on a digitally mounted complete skeleton. Palaeontologia Electronica 13 (2): 1–26.
Mallison H., Hohloch A. & Pfretzschner H.-U. 2009. Mechanical digitizing for paleontology – new and improved techniques. Palaeontologia Electronica 12 (2) 4T: 1–41.
Mantell G.A. 1825. Notice on the Iguanodon, a newly discovered fossil reptile, from the sandstone of Tilgate Forest, in Sussex. Philosophical Transactions of the Royal Society of London 115: 179–186. https://doi.org/10.1098/rstl.1825.0010
Mantell G.A. 1833. The Geology of the South-East of England. First Edition. Longman, Rees, Orme, Brown, Green, & Longman, London. https://doi.org/10.5962/bhl.title.106921
Martínez R.N. 2009. Adeopapposaurus mognai, gen. et sp. nov. (Dinosauria: Sauropodomorpha), with comments on adaptations of basal Sauropodomorpha. Journal of Vertebrate Paleontology 29 (1): 142–164. https://doi.org/10.1671/039.029.0102
Marzola M., Mateus O., Milàn J. & Clemmensen L. 2018. A review of Palaeozoic and Mesozoic tetrapods from Greenland. Bulletin of the Geological Society of Denmark 66: 21–46. https://doi.org/10.37570/bgsd-2018-66-02
Mayr E. 1968. The role of systematics in biology: the study of all aspects of the diversity of life is one of the most important concerns in biology. Science 159 (3815): 595–599. https://doi.org/10.1126/science.159.3815.595
Mones A. 1989. Nomen dubium vs. nomen vanuum. Journal of Vertebrate Paleontology 9 (2): 232–234. https://doi.org/10.1080/02724634.1989.10011757
Moser M. 2003. Plateosaurus engelhardti Meyer, 1837 (Dinosauria: Sauropodomorpha) from the Feuerletten (Middle Keuper; Upper Triassic) of Bavaria. Zitteliana B (24): 3–186. https://doi.org/10.5282/ubm/epub.12711
Nitsch E. & Hagdorn H. 2015. Lettenkohle, Lettenkeuper, Unterkeuper – Vom Gesteinsnamen zum stratigraphischen Begriff. Palaeodiversity Suppl. 1: 17–23.
Owen R. 1842. Report on British fossil reptiles. Report of the Eleventh Meeting of the British Association for the Advancement of Science; Held at Plymouth in July 1841 II: 60–205.
Plieninger T. 1846a. Über ein neues Sauriergenus und die Einreihung der Saurier mit flachen, schneidenden Zähnen in eine Familie. Jahreshefte des Vereins für vaterländische Naturkunde in Württemberg 2: 148–154.
Plieninger T. 1846b. Nachträgliche Bemerkungen zu dem Vortrage (S. 148 dieses Heftes) über ein neues Sauriergenus und die Einreihung der Saurier mit flachen, schneidenden Zähnen in eine Familie. Jahreshefte des Vereins für vaterländische Naturkunde in Württemberg 2: 247–254.
Plieninger T. 1847. II. Aufsätze und Abhandlungen. 1. Verzeichniss der Reptilien Württembergs. Jahreshefte des Vereins für vaterländische Naturkunde in Württemberg 3: 194–208.
Plieninger T. 1852. Belodon plieningeri H. v. Meyer, Ein Saurier der Keuperformation. Jahreshefte des Vereins für vaterländische Naturkunde in Württemberg 8: 389–524.
Prieto-Márquez A. & Norell M.A. 2011. Redescription of a nearly complete skull of Plateosaurus (Dinosauria: Sauropodomorpha) from the Late Triassic of Trossingen (Germany). American Museum Novitates 3727: 1–58. https://doi.org/10.1206/3727.2
Quenstedt F.A. 1852. Handbuch der Petrefaktenkunde. H. Laupp’sche Buchhandlung, , Tübingen.
Quenstedt F.A. 1855. Über Pterodactylus suevicus im lithographischen Schiefer Württembergs. H. Laupp’sche Buchhandlung, Tübingen.
Quenstedt F.A. 1867. Handbuch der Petrefaktenkunde. H. Laupp’sche Buchhandlung, Tübingen.
Quenstedt F.A. 1885. Handbuch der Petrefaktenkunde. H. Laupp’sche Buchhandlung, Tübingen. https://doi.org/10.5962/bhl.title.30642
Rauhut O.W.M., Holwerda F.M. & Furrer H. 2020. A derived sauropodiform dinosaur and other sauropodomorph material from the Late Triassic of Canton Schaffhausen, Switzerland. Swiss Journal of Geosciences 113 (8): 1–54. https://doi.org/10.1186/s00015-020-00360-8
Regalado Fernández O.R. & Werneburg I. 2022. A new massopodan sauropodomorph from Trossingen Formation (Germany) hidden as ‘Plateosaurus’ for 100 years in the historical Tübingen collection. Vertebrate Zoology 72: 771–822. https://doi.org/10.3897/vz.72.e86348
Reinacher S.J. 2021. 1921 bis 1923: Plateosaurus engelhardti – zwei schwäbische Lindwürmer. In: Seidl E., Bende E. & Werneburg I. (eds) Aus der Tiefenzeit: Die Paläontologische Sammlung der Universität Tübingen. First Edition: 394–411. Museum der Universität MUT, Tübingen.
Rieß-Stumm S. 2021. Anno 1922 Dokumentation zur Dinosauriergrabung in Trossingen. In: Werneburg I. (ed.) Chelyops. Berichte aus der Paläontologischen Sammlung in Tübingen: 116–118. Scidinge Hall Verlag, Tübingen.
Riley H. & Stutchbury S. 1836. A description of various fossil remains of three distinct Saurian animals discovered in the autumn of 1834, in the Magnesian Conglomerate on Durham Down, near Bristol. Proceedings of the Geological Society of London 2 (11): 397–399.
Rütimeyer L. 1856. Fossile Reptilienknochen aus dem Keuper. Verhandlungen der Schweizerischen Naturforschenden Gesellschaft 41: 62–64.
Sander P.M. 1992. The norian Plateosaurus bonebeds of central-Europe and their taphonomy. Palaeogeography, Palaeoclimatology, Palaeoecology 93: 255–299. https://doi.org/10.1016/0031-0182(92)90100-J
Sander P.M. & Klein N. 2005. Developmental plasticity in the life history of a prosauropod dinosaur. Science 310 (5755): 1800–1802. https://doi.org/10.1126/science.1120125
Schaffer J. & Schoch R.R. 2022. Redescription of the neotype of Plateosaurus. In: Schweigert G. (ed.) Abstract Volume. 93. Jahrestagung der Paläontologischen Gesellschaft, Sep. 19th to 23rd at the Staatliches Museum für Naturkunde Stuttgart: 60. Naturkunde Museum Stuttgart, Stuttgart.
Schoch R.R. 2011. Tracing Seemann’s dinosaur excavation in the Upper Triassic of Trossingen: his field notes and the present status of the material. Palaeodiversity 4 (December): 245–282.
Seidl E., Bierende E. & Werneburg I. 2021. Aus der Tiefenzeit. Die Paläontologische Sammlung der Universität Tübingen. Publikationen des Museums der Universität Tübingen, Tübingen.
Steel R. 1970. Handbuch der Paläoherpetologie. Part 14: Saurischia. Gustav Fischer, Stuttgart.
Steyskal G.C. 1965. Trend curves of the rate of species description in zoology. Science 149 (3686): 880–882. https://doi.org/10.1126/science.149.3686.880
Sues H.-D. 2013. Comment on Case 3560: Plateosaurus engelhardti Meyer, 1837 (Dinosauria, Sauropodomorpha): proposed replacement of unidentifiable name-bearing type by a neotype (Case 3560; see BZN 69: 203–212, 295–296). Bulletin of Zoological Nomenclature 70 (2): 120–121.
Turrill W.B. 1938. The expansion of taxonomy with special reference to Spermatophyta. Biological Reviews 13 (4): 342–373. https://doi.org/10.1111/j.1469-185X.1938.tb00522.x
Urlichs M. 1966. Zur Fossilführung und Genese des Feuerlettens, der Rhät-Lias-Grenzschichten und des unteren Lias bei Nürnberg. Erlanger Geologische Abhandlungen 64: 42.
von Huene F.F. 1905. Trias-Dinosaurier Europas. Zeitschrift der Deutschen Geologischen Gesellschaft 57: 345–349.
von Huene F.F. 1907. Die Dinosaurier der europäischen Triasformation mit Berücksichtigung der aussereuropäischen Vorkommnisse. First Edition. Gustav Fischer, Tübingen.
von Huene F.F. 1908. Eine Zusammenstellung über die englische Trias und das Alter ihrer Fossilien. Centralblatt für Mineralogie, Geologie und Paläontologie, Jahrgang 1908 (1): 9–17.
von Huene F.F. 1914. Saurischia et Ornithischia triadica (‘Dinosauria’ triadica). W. Junk, Berlin.
von Huene F.F. 1915. Beiträge zur Kenntnis einiger Saurischier der schwäbischen Trias. Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 1915: 1–27.
von Huene F.F. 1926. Vollständige Osteologie eines Plateosauriden aus dem schwäbischen Keuper. Geologische und Palaeontologische Abhandlungen, Neue Folge 15 (2): 139–179.
von Huene F.F. 1928. Lebensbild des Saurischier-Vorkommens im Obersten Keuper von Trossingen in Württemberg. Paleobiologica 1: 103–116.
von Huene F.F. 1929. Kurze Überischt über die Saurischia und ihre natürlichen Zusammenhänge. Paläontologische Zeitschrift 11: 269–273. https://doi.org/10.1007/BF03042732
von Huene F.F. 1932. Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte. Monographien zur Geologie und Paläontologie 2 (4): 1–192.
von Huene F.F. 1959. Paläontologie und Phylogenie der niederen Tetrapoden: Nachträge und Ergänzungen. Fischer, Jena.
von Koken E. 1904. Das geologisch-mineralogische Institut in Tübingen. Centralblatt für Mineralogie, Geologie und Paläontologie 1904: 673–693.
von Meyer H. 1832. Palaeologica zur Geschichte der Erde und ihrer Geschöpfe. Verlag von Siegmund Schmerber, Frankfurt am Main.
von Meyer H. 1837. Mittheilungen, an Professor Bronn gerichtet vom 4. April 1837. Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde 1837: 314–316.
von Meyer H. 1839. Mittheilungen, an Professor Bronn gerichtet vom 1. Dezember 1838. Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde: 76–79.
von Meyer H. 1855. Zur Fauna der Vorwelt. II. Die Saurier des Muschelkalkes mit Rücksicht auf die Saurier aus Buntem Sandstein und Keuper. S. Schmerber’schen Buchhandlung, Frankfurt am Main.
von Meyer. H. 1861. Reptilien aus dem Stubensandstein des oberen Keupers. Palaeontographica 7: 253–346.
Wagner J.A. 1851. Beiträge zur Kenntniss der im süddeutschen Lias vorkommenden Arten von Ichthyosaurus. Abhandlungen der königlich Bayerischen Akademie der Wissenschaften 2. Classe 6 (2): 485–538.
Walker A.D. 1964. Triassic reptiles from the Elgin area: Ornithosuchus and the origin of carnosaurs. Philosophical Transactions of the Royal Society B: Biological Sciences 248 (744): 53–134. https://doi.org/10.1098/rstb.1964.0009
Wang Y.-M., Wang T., Yang Z.W. & You H.L. 2019. Cranium and vertebral column of Xingxiulong chengi (Dinosauria: Sauropodomorpha) from the Early Jurassic of China. Anatomical Record 303 (4: The Hidden World of Dinosaurs): 772–789. https://doi.org/10.1002/ar.24305
Weishampel D.B. 1984. E. Fraas, F. von Huene, R. Seemann, and the ‘Schwabische Lindwurm’ Plateosaurus. In: Reif W.-E. & Westphal F. (eds) Third Symposium on Mesozoic Terrestrial Ecosystems, short papers: 249–253. Attempto Verlag, Tübingen.
Weishampel D.B. & Chapman R.E. 1990. 3 – Morphometric study of Plateosaurus from Trossingen (Baden-Wurttemberg, Federal Republic of Germany). In: Carpenter K. & Currie P.J. (eds) Dinosaur Systematics: 43–52. Cambridge University Press, Cambridge. https://doi.org/10.1017/CBO9780511608377.006
Weishampel D.B. & Westphal F. 1986. Die Plateosaurier von Trossingen. Ausstellungs-Kataloge University Tübingen 19: 1–27.
Wellnhofer P. 1993. Prosauropod dinosaurs from the Feuerletten (Middle Norian) of Ellingen near Weissenburg in Bavaria. Revue de Paléobiologie. Volume spécial 7: 265–271.
Werneburg I. & Böhme M. 2018. TÜBINGEN: The Palaeontologial Collection of Tübingen. In: Beck L.A. & Joger U. (eds) Paleontological Collections of Germany, Austria and Switzerland. First Edition: 505–512. Springer, Cham. https://doi.org/10.1007/978-3-319-77401-5_52
Yates A.M. 2003. The species taxonomy of the sauropodomorph dinosaurs from the Löwenstein Formation (Norian, Late Triassic) of Germany. Palaeontology 46 (2): 317–337. https://doi.org/10.1111/j.0031-0239.2003.00301.x
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