A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)

Henrik Enghoff, Sara B. Frederiksen


The genus Aquattuor Frederiksen, 2013 is revised. A. denticulatus Frederiksen, 2013 (type species) from the East Usambara Mts, Tanzania, is redescribed, and six new species are described: A. claudiahempae Enghoff & Frederiksen sp. nov. from Mt. Kilimanjaro, Tanzania, and five species from the Udzungwa Mts., Tanzania: A. longipala Enghoff sp. nov., A. major Enghoff sp. nov., A. stereosathe Enghoff sp. nov., A. submajor Enghoff sp. nov., and A. udzungwensis Enghoff sp. nov.


Eastern Arc; Kilimanjaro; taxonomy; new species

Full Text:


DOI: https://doi.org/10.5852/ejt.2015.150


Akkari N. & Enghoff H. 2011. On some surface structures of potential taxonomic importance in families of the suborders Polydesmidea and Dalodesmidea (Polydesmida, Diplopoda). Zookeys 156: 1–24. http://dx.doi.org/10.3897/zookeys.156.2134

Attems C. 1909. Myriopoda. Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905-1906 3 (19): 1–64.

Burgess N., Butinsky T.M., Cordeiro N.J., Doggart N.H., Fjeldså J., Howell K.M., Kilahama F.B., Loader S.P., Lovett J.C., Mbilinyi B., Menegon M., Moyer D.C., Nashanda E., Perkin A., Rovero F., Stanley W.T. & Stuart S.N. 2007. The biological importance of the Eastern Arc Mountains of Tanzania and Kenya. Biological Conservation 134: 209–231. http://dx.doi.org/10.1016/j.biocon.2006.08.015

Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1–75. http://dx.doi.org/10.5852/ejt.2014.100

Frederiksen S.B. 2013a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597–600. http://dx.doi.org/10.11646/zootaxa.3636.4.7

Frederiksen S.B. 2013b. East African odontopygid millipedes 3: Two new genera; Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59–66. http://dx.doi.org/10.11646/zootaxa.3694.1.4

Frederiksen S.B. & Enghoff H. 2012. East African odontopygid millipedes 1: Five new species of Xystopyge (Attems, 1909) and a proposal for a new gonopod terminology (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3485: 69–82.

Frederiksen S.B. & Enghoff H. 2015. East African odontopygid millipedes 4: A restricted redefinition of the genus Rhamphidarpoides Kraus, 1960, a related new genus, five new species, and notes on solenomere function (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3926: 541–560. http://dx.doi.org/10.11646/zootaxa.3926.4.5

Kraus O. 1960. Äthiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1–207.

Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 512: 1–143.

Marshall A.R., Jørgensbye H.I.O., Rovero F., Platts P.L., White P.C.L. & Lovett J.C. 2010. The species–area relationship and confounding variables in a threatened monkey community. American Journal of Primatology 72: 325–336. http://dx.doi.org/10.1002/ajp.20787

Nonnotte P., Guillou H., Le Gall B., Benoit M., Cotten J. & Scaillet S. 2008. New K–Ar age determinations of Kilimanjaro volcano in the North Tanzanian diverging rift, East Africa. Journal of Volcanology and Geothermal Research 173 (1–2): 99–112. http://dx.doi.org/10.1016/j.jvolgeores.2007.12.042

Reboleira A.S.P.S. & Enghoff H. 2015. Redescription of Lusitanipus alternans (Verhoeff, 1893) (Diplopoda, Callipoda, Dorypetalidae) and ecological data on its Laboulbeniales ectoparasites in caves. Zootaxa 3957: 567–576. http://dx.doi.org/10.11646/zootaxa.3957.5.5

Soini E. 2005. Changing livelihoods on the slopes of Mt. Kilimanjaro, Tanzania: Challenges and opportunities in the Chagga homegarden system. Agroforestry Systems 64: 157–167. http://dx.doi.org/10.1007/s10457-004-1023-y


  • There are currently no refbacks.

License URL: https://creativecommons.org/licenses/by/3.0/