New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia)
Abstract
The brachiopods Meristorygma donakovae Poletaev, 2018, Johnsoniana nom. nov. from Mississippian strata of the Urals and Betaneospirifer stepanovi Poletaev, 2015 from Gzhelian rocks of the Urals are re-described. Tiramnia davidi sp. nov. from the Bashkirian of the Donets Basin (Ukraine) is newly described. Revision of available data shows that the revised genus Verneuilia Hall & Clarke, 1893 belongs to the family Elythynidae Gourvennec, 1994 of the superfamily Martinioidea Waagen, 1883. The genera Johnsoniana nom. nov., Changshunella Sun et al., 2004 and possibly Minythyra Branton, 1984 belong to the family Oceaniidae Poletaev, 2015 of the superfamily Paeckelmanelloidea Ivanova, 1972. The importance of characters such as denticulation of the hinge line of the interarea and type of vascular impression in the diagnosis of higher spiriferid taxa is shown.
References
Aisenverg D.E. 1950. Materialy k faune brakhiopod svity C23 Donetskogo basseyna. Materialy po stratigrafii i paleontologii Donetskogo basseyna: 113–141. Moskow, Ugletekhizdat.
Aisenverg D.E. 1951. Brakhiopody kamennougol’nykh otlozheniy rayona reki Volch’yey. Trudy Instituta geologicheskich nauk Akademii Nauk Ukrainckoy SSR. Seriya stratigrafiya i paleontologiya 5: 1–49.
Aisenverg D.E. & Poletaev V.I. 1971. Brachiopody. In: Aisenverg D.E. (ed.) Atlas fauny turneyskich otlozheniy Donetskogo basseyna (s opisaniyem novych vidov): 66–91. Naukova Dumka, Kiev.
Aisenverg D.E. & Poletaev V.I. 1983. Brachiopody. In: Didkovskiy V.Ya. (ed.) Verkhneserpukhovskoy pod’yarus Donetskogo Basseyna (paleontologicheskaya kharakteristika): 73–95. Naukova Dumka, Kiev.
Aleksandrov V.A. & Einor O.L. 1979. Brakhiopody. In: Einor O.L. (ed.) Atlas fauny i flory srednego-pozdnego karbona Bashkirii: 55–97. Nedra, Moscow.
Aleksandrov V.A. & Einor O.L .1996. Middle Carboniferous brachiopods, mainly from the Bashkirian of the South Urals (stratotype area). In: Wagner R.H., Winkler-Prins C.F. & Granados L.F. (eds) The Carboniferous of the World. The Former USSR, Mongolia, Middle Eastern Platform, Afghanistan & Iran 3: 306–307. Instituto Tecnológico Geominero de España, Madrid.
Angiolini L., Long S. & Davies L. 2011. Revision of Sowerby’s species Spirifer bisulcatus, Spirifer pinguis and Spirifer rotundatus from the late Tournaisian–Visean of Great Britain. Memoirs of the Association of Australasian Palaeontologists 41: 71–85.
Angiolini L., Cisterna G.A., Mottequin B., Shen S.-Z. & Muttoni G. 2022. Global Carboniferous brachiopod biostratigraphy. Special Publication of the Geological Society 512 (1): 497–550. https://doi.org/10.1144/sp512-2020-225
Archbold N.W. & Thomas G.A. 1984. Neospirifer Frederiks, 1924 (Spiriferida, Brachiopoda): A review. Journal of Paleontology 58 (3): 626–635.
d’Archiac V.E. & de Verneuil E.P. 1842. On the fossils of the older deposits in the Rheinish Provinces. Transactions Geological Society 6 (2): 303–410. https://doi.org/10.1144/transgslb.6.2.303
Barkhatova V.P. 1970. Biostratigrafiya nizhnego karbona i permi Severnogo Timana. Trudy Vsesoyuznogo Geologo-razvedochnogo Instituta 283: 1–229.
Beznosova G.A. 1968. Otryad Spiriferida. In: Sarycheva T.G. (ed.) Brakhiopody verkhnego paleozoya Vostochnogo Kazakhstana. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 121: 166–200.
Blodgett R.B. & Johnson J.G. 1994. First recognition of the genus Verneuilia Hall and Clarke (Brachiopoda, Spiriferida) from North America (West-Central Alaska). Journal of Paleontology 68 (6): 1240–1242. https://doi.org/10.1017/S0022336000034235
Brunton C.H.C. 1984. Silicified brachiopods from the Viséan of County Fermanagh, Ireland. (III). Rhynchonellids, spiriferids and terebratulids. Bulletin of the British Museum (Natural History) Geology 38 (2): 27–127. Available from https://www.biodiversitylibrary.org/page/2273038 [accessed 7 Oct. 2024].
Carniti A.P., Della Porta G., Banks V.J., Stephenson M.H. & Angiolini L. 2022. Brachiopod fauna from uppermost Viséan (Mississippian) mud mounds in Derbyshire, UK. Acta Palaeontologica Polonica 67 (4): 865–915. https://doi.org/10.4202/app.00972.2022
Carter J.L. 1974. New genera of spiriferid and brachythyridid brachiopods. Journal of Paleontology 48 (4): 674–696.
Carter J.L. & Gourvennec R. 2007. Spiriferida and Spiriferinida. In: Selden P. (ed.) Treatise on Invertebrate Paleontology, Pt H, Revised Brachiopoda 6: 2772–2796. The Geological Society of America, The University of Kansas, Boulder and Lawrence.
Carter J.L. & Poletaev V.I. 1998. Atokan (Late Bashkirian or Early Moscovian) brachiopods from the Hare Fiord formation of Ellesmere Island, Canadian Arctic Archipelago. Annals of Carnegie Museum 67 (2): 105–180. https://doi.org/10.5962/p.215207
Carter J.L., Johnson J.G., Gourvennec R. & Hongfei H. 1994. A revised classification of the spiriferid brachiopods. Annals of the Carnegie Museum 63: 327–374.
Available from https://www.biodiversitylibrary.org/part/215817 [accessed 10 Oct. 2024]
Carter J.L., Johnson J.G., Gourvennec R. & Hou H.-f. 2006. Spiriferida. In: Moore R.C. (ed.) Treatise on Invertebrate Paleontology. Pt H, Revised Brachiopoda 5: 1689–1937. The Geological Society of America, The University of Kansas, Lawrence.
Chao Y.T. 1929. Carboniferous and Permian Spiriferids of China. Palaeontologia Sinica 11: 1–101.
Chernyshev F.N. 1902. Verkhnekamennougol’nyye brakhiopody Urala i Timana. Trudy Geologicheskogo komiteta 16: 1–432.
Davidson T. 1858–1863. A Monograph of the British Fossil Brachiopoda. Vol. 2, рt 5. London. https://doi.org/10.1017/cbo9781139104067.002
Donakova L.M. 1983. O vozmozhnosti vydeleniya kosogorskogo i brazhkinskogo gorizontov nizhnego karbona na vostochnom sklone Yuzhnogo Urala. Yezhegodnik Vsesoyuznogo paleontologicheskogo obshchestva 26: 207–227.
Einor O.L. 1936. Stratigrafiya i rukovodyashchaya brachiopodovaya fauna izvestnyakov naduglenosnoy tolshchy nizhnrgo karbona Kizelovskogo rayona na Urale: 1–32. ОNТY NKTP SSSR.
Einor O.L. 1957. К faune kamennougol’nych brachiopod Zapadnogo sklona Srednego Urala. Yezhegodnik Vsesoyuznogo paleontologicheskogo obshchestva 16: 142–161.
Eschscholtz F. 1829. System der Acalephen. Eine ausführliche Beschreibung aller medusenartigen Strahlthiere. Ferdinand Dümmler, Berlin. https://doi.org/10.5962/bhl.title.64070
Frederiks G.N. 1911. Bemerkung über einige Oberpaläeozoische Fosilien von Krasnoufimsk. Prilozhenie k Protokolam Zasedaniy Obshchestva Estestvoispytateley pri Imperatorskom Kazanskom Universitete 42 (269): 1–11.
Frederiks G.N. 1924. Paleontologicheskiye etyudy. 2. O verkhnekamennougol’nykh spiriferidakh Urala. Izvestiya Geologicheskogo Komiteta 38 (3): 295–324.
Gatinaud G. 1949. Contributions à l’étude des brachiopodes Spiriferidae. 1. Exposé d’une nouvelle méthode d’étude de la morphologie externe des Spiriferidae à sinus plissé. Bulletin du Muséum national d’Histoire naturelle (2e Série) 21: 153–492. Available from https://www.biodiversitylibrary.org/page/54382725 [accessed 7 Oct. 2024].
George T.N. 1931. Ambocoelia Hall and certain similar British Spiriferidae. Quarterly Journal of the Geological Society of London 87: 30–61. https://doi.org/10.1144/gsl.jgs.1931.087.01-04.05
Gerasimov N.P. 1929. Brakhiopody sterlitamakskogo izvestnyaka. Uchonyye zapiski Kazanskogo universiteta 39 (5/6): 779–872.
Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573–581. https://doi.org/10.1016/s0016-6995(94)80253-x
Grunt T.A. 1977. Kamennougol’nyye spiriferidy semeystva Martiniidae Sibiri i Arktiki. In: Brakhiopody verkhnego paleozoya Sibiri i Arktiki. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 162: 63–71.
Hall J. & Clarke J.M. 1893. An Introduction to the Study of the Genera of Paleozoic Brachiopoda: intended as a Handbook for the Use of Students. Palaeontology of New York 8 Pt. 2. James B. Lyon, Albany. https://doi.org/10.5962/bhl.title.15111
ICZN (International Commission on Zoological Nomenclature) 1999. International Code of Zoological Nomenclature. 4th edition. International Trust for Zoological Nomenclature and The Natural History Museum, London. Available from https://www.iczn.org/the-code/the-code-online/ [accessed 7 Oct. 2024].
Ilovajsky D. 1929 Novye dannye po plechenogim iz svit М i N Donetskogo Baseyna. Izvestiya Geologichaskogo Komiteta 48 (2): 83–97.
Ivanova E.A. 1960. Otriad Spiriferida. In: Orlov Y.A. (ed.) Osnovy Paleontologii. Mshanki, Brachiopody. Vol. 7: 264–280. Izdatel’stvo Akademii Nauk SSSR, Moscow.
Ivanova E.A. 1972. Oshovnye zakonomernosty evolyutsyi spiriferid (Brachiopoda). Paleontologichesky Zhurnal 3: 28–42.
Kalashnikov N.V. 1974. Rannekamennougol’nye brachiopody Pechorskogo Urala. Nauka, Leningrad.
Kalashnikov N.V. 1980. Brakhiopody verkhnego paleozoya Yevropeiskogo Severa SSSR. Nauka, Leningrad.
Keyserling A. de. 1846. Geognostische Beobchtungen. In: Wissenschaftliche Beobachtungen auf einer Reise in das Petschora−Land im Jahre 1843: 149–406. Carl Kray, St. Petersburg.
dе Koninck L.G. 1842–1844. Description des Animaux fossiles qui se trouvent dans le Terrain carbonifére de Belgique. H. Dessain, Liège. Available from https://www.biodiversitylibrary.org/bibliography/197887 [accessed 7 Oct. 2024].
de Koninck, L.G. 1883. Sur le Spirifer Mosquensis et sur ses affinités avec quelques autres espèces du même genre. Bulletin du Musée royal d’Histoire naturelle de Belgique 2: 371-395.
dе Koninck L.G. 1887. Faune du Calcaire Carbonifère de la Belgique. Brachiopodes. Annales du Musée royal d’histoire naturelle de Belgique 14: 1–159. https://doi.org/10.5962/bhl.title.149304
Kulagina E.I., Pazukhin V.N. & Davydov V.I. 2009. Pennsylvanian biostratigraphy of the Basu River Section with emphasis on the Bashkirian–Moscovian transition. In: Puchkov V.N. (ed.) Carboniferous Type Sections of Russia and Potential Global Stratotypes: 42–63. Ufa.
Kutorga S.S. 1844. Zweiter Beiträge zur Palaeontologie Russlands. Verhandlungen der Russisch-Kaiserlichen mineralogischen Gesellschaft zu St. Petersburg 1844: 62–104.
Lapina N.N. 1975. Brakhiopody. In: Stepanov D.L., Krylova A.K., Grozdilova L.P., Pozner V.M. & Sultanaev A.A. (eds) Paleontologicheskiy atlas kamennougol’nykh otlozheniy Urala. Trudy Vsesoyuznogo nauchno-issledovatel’skogo geologo-razvedochnogo instituta 383: 154–203.
Lazarev S.S. & Poletaev V.I. 1982. The development of the Brachiopod mantle canal system at the early-middle Carboniferous boundary. In: Ramsbottom W.H. (ed.) Biostratigraphic Data for a Mid-Carboniferous Boundary: 89–94. Leeds.
Lebedev N.I. 1913. Spiriferidae iz kamennougol’nych otlozheniy Donetskogo basseyna I nekotorych drugich rayonov Rossii. Materialy dlya geologii Donetskogo kamennougol’nogo basseyna. Izvestiya Ekaterinoslavskogo gornogo instituta 1: 1–21.
Lі L. & Gu F. 1980. Late Carboniferous brachiopods from Yanji of Jilin, northeast China. Acta Paleontologica Sinica 19 (6): 483–491.
Licharew B.K. 1938. Materialy do vyvchenn’ya verchn’okam’yanovugil’nych brachiopod Donets’kogo baseynu. Geologichny Zhurnal 5/4 (12): 73–129.
Lisitsyn K.I. 1925. Podrazdeleniya nizhnego karbona i ikh korallovo-brachiopodovaya fauna. Izvestiya Donskogo politekhnicheskogo instituta 13: 1–117.
M’Coy F. 1844. A Synopsis of the Characters of the Carboniferous Limestone Fossils of Ireland. Williams & Nergate, London. https://doi.org/10.5962/bhl.title.11559
Mironova M.G. 1967. Pozdnekamennougol’nyye brakhiopody Bashkirii. Izdatel’stvo Leningradskogo universiteta, Leningrad.
Mottequin B. & Poty E. 2022. Brachiopods from the historical type area of the Viséan Stage (Carboniferous, Mississippian; Belgium) and the Visé fauna: preliminary remarks. Palaeobiodiversity and Palaeoenvironments 102: 351–371. https://doi.org/10.1007/s12549-021-00498-9
Nalivkin D.V. 1979. Brakhiopody turneyskogo yarusa Urala. Nauka, Leningrad.
Nemyrovska T.I. & Yefimenko V.І. 2013. Seredniy karbon (nyzhniy pensylvaniy). In: Gozhyk P.F. (ed.) Stratigraphy of the Upper Proterozoic and Phanerozoic of Ukraine. Vol. 1. Stratigraphy of the Upper Proterozoic, Paleozoic and Mesozoic of Ukraine: 283–303. Logos, Kyiv.
d’Orbigny A. 1850. Mémoire sur les Brachiopodes. II. Classification des Brachiopodes. Annales des Sciences Naturelles. Zoologie, Série 3e 13: 295–353. Available from https://www.biodiversitylibrary.org/page/13434796 [accessed 7 Oct. 2024].
Péron F. & Lesueur C.A. 1810. Tableau des caractères génériques et spécifiques de toutes les espèces de méduses conneus jusqu’à ce jour. Annales du Muséum d’Histoire naturelle 14: 325–366. Available from https://www.biodiversitylibrary.org/page/3498981 [accessed 7 Oct. 2024].
Pitrat C.W. 1965. Spiriferidina. In: Moore R.C. (ed.) Treatise on Invertebrate Paleontology, Part H, Brachiopoda: H667–H728. Geological Society of America and University of Kansas, Lawrence.
Poletaev V.I. 1975. Rannekamennougol’nye i bashkirskie gladkie spiriferidy i atiridy Donetskogo basseyna. Naukova Dumka, Кiev.
Poletaev V.I. 1997. Revision of the genus Neospirifer Frederiks, 1924 and the new genus from subfamily Neospiriferinae – Lutuginia. Paleontological Journal 3: 54–63.
Poletaev V.I. 1999. The revision of Palaeochoristites and the new genus of the Lower Carboniferous spiriferide, Phragmobrachythyris. Paleontological Journal 3: 41–49.
Poletaev V.I. 2008. Brachiopody semeistva Martiniidae Waagen, 1883 iz srednego karbona Donetskogo basseyna. In: Gozhyk P.F. (ed.) Bíostratigrafíchní osnovy pobudovy stratygrafíchnykh skhem fanerozoyu Ukraíny. Zbirnyk naukovykh prats’ Ínstututu geologichnykh nauk Natsional’noí Akademií Nauk Ukraíny: 27–35.
Poletaev V.I. 2009. Novye dannye o nomenklature podsemeistva Angiospiriferinae Legrand-Blain, 1985 (brachiopody karbona). In: Gozhyk P.F. (ed.) Vykopna fauna í flora Ukrainy: paleoyekologíchniy ta stratigrafíchniy aspekty. Zbirnyk naukovykh prats Ínstututu geologichnykh nauk Natsionalnoí Akademií Nauk Ukraíny: 69–72.
Poletaev V.I. 2013. Osnovnyye etapy evolyutsii vnutrennego stroyeniya nekotorykh rodov kamennougol’nykh spiriferid Paleotetisa. Zbirnyk naukovykh prats Ínstututu geologichnykh nauk Natsionalnoí Akademií Nauk Ukraíny 6 (1): 11–16. https://doi.org/10.30836/igs.2522-9753.2009.147894
Poletaev V.I. 2015. Novyy rod i semeystvo kamennougol’nykh spiriferid Yevropy. In: Gozhyk P.F. (ed.) Stratotypoví ta oporní rozrízy fanerozois’kykh vídkladív Ukraíny: suchasnyi stan ta perspektyvy podal’shykh doslídzhen’. Materíaly 36 sesií Paleontologichnogo tovarustva Ukraíny: 17–18. Kyiv.
Poletaev V.I. 2017. Novi dani do systematyky i nomenklatury kamʹyanovuhilʹnykh spiryferyd Yevropy. In: Gozhyk P.F. (ed.) 40 rokív Paleontologíchnomu Tovaristvu Ukraíny. Materíaly 38 sesií Paleontologichnogo tovarustva Ukrainy: 74. Kyiv.
Poletaev V.I. 2018. Atlas-opredelitel’ kamennougol’nykh spiriferid Vostochnoy Yevropy. LAT&K, Kyiv.
Poletaev V.I. 2022. On the history of systematics and the evolution of the brachiopods order Spiriferida from the Carboniferous of Eastern Europe. Paleontological Journal 56 (7): 118–129. https://doi.org/10.1134/s0031030122070127
Quenstedt F.A. 1868–1871. Petrefactenkunde Deutschlands. 1. Abt., 2. Band: Die Brachiopoden. Tubingen & Leipzig. 1–160 (1868); 161–464 (1869); 465–748 (1870); 25 pl. (1871).
Reed F.R.C. 1944. Brachiopods and Mollusca from the Productus Limestones of the Salt Range. Palaeontologica Indica (new series) 23 (2): 1–678.
Rotai A.P. 1931. Brakhiopody i stratigrafiya nizhnego karbona Donetskogo basseyna. Trudy Glavnogo Geologo-Razvedochnogo Upravleniya 73: 35–117.
Rotai A.P. 1951. Brakhiopody srednego karbona Donetskogo basseyna. 1. Spiriferidae. Gosgeolizdat, Moscow.
Schuchert C. 1893. A classification of the Brachiopoda. The American Geologist 11 (3): 141–167. Available from https://www.biodiversitylibrary.org/page/54930209 [accessed 7 Oct. 2024].
Schuchert C. & LeVene C.M. 1929. Brachiopoda (Generum et Genotyporum Index et Bibliographia). In: Pompeckj J.F. (ed.) Fossilium Catalogus. I: Animalia Pars 42: 1–140. W. Junk, Berlin.
Semikhatova S.V. 1941a. Brachiopody bashkirskikh sloyev SSSR. 1. Rod Choristites Fischer. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 12 (4): 1–152.
Semikhatova S.V. 1941b. Gruppa «Spirifer trigonalis Martin» v naduglenosnykh sloyakh nizhnego karbona Podmoskovnogo basseyna. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 12 (3): 5–175.
Stepanov D.L. 1948. Verkhnekamennougol’nyye brakhiopody Bashkirii. Trudy Vsesoyuznogo nauchno-issledovatel’skogo geologo-razvedochnogo instituta, novaya seriya 22: 3–64.
Sun Y., Baliński A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267–274.
Tiazheva A.P. 1960. Novyy vid devonskikh retikulyariid Urala. In: Markovskiy B.P. (ed.) Novyye vidy drevnikh rasteniy i bespozvonochnykh SSSR: 406–409. Vsecoyuznyi geologicheskiy institute, Leningrad.
Waagen W.H. 1883. Salt Range fossils, vol. 1, part 4. Productus Limestone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, series 13 2: 391–546.
Wang C. & Yang S. 1993. Brachiopod fauna around Carboniferous–Permian boundary from Balikelike Formation in Keping, Xinjiang. Journal of Changchun University of Geosciences 23 (1): 6–9.
Wang Y., Jin Y.G. & Fang D.W. 1966. Fossil Brachiopoda. Science Press, Beijing.
Wang Y.N., Ma X.P., Ebbighausen V. & Becker R.T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Paläontologische Zeitschrift 97: 19–35. https://doi.org/10.1007/s12542-022-00635-5
Waterhouse J.B. 1968. The classification and description of Permian Spiriferida (Brachiopoda) from New Zealand. Palaeontographica 129 (1–3): 1–94.
Waterhouse J.B. 2004. Permian and Triassic stratigraphy and fossils of the Himalaya in Northern Nepal. Earthwise 6: 1–265.
Yanishevsky M.E. 1900. Fauna kamennougol’nogo izvestnyaka, vystupayushchego po r. Shartymke, na vostochnom sklone Urala. Trudy Obshchestva yestestvoispytateley pri Imperatorskom Kazanskom universitete 34 (5): 1–140.
Yanishevsky M.E. 1910. Nizhnekamennougol’nyy izvestnyak okolo Khabarnogo, Orskogo uyezda Orenburgskoy gubernii. Izvestiya Tomskogo politechnicheskogo instituta 17 (1): 1–305.
Yanishevsky M.E. 1918. Materialy k izucheniyu nizhnekamennougol’noy fauny Fergany. Trudy Geologicheskogo Komiteta, novaya seriya 162: 1–145.
Zhang C., Zhang F.M., Zhang Z.X. & Wang Z. 1983. Phylum Brachiopoda. In: Paleontological Atlas of Northwestern China for Xinjiang Uygur Autonomous Region, Volume 2 (Upper Paleozoic): 262–386. Geological Publishing House, Beijing.
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