New data on zodariid spiders (Arachnida, Araneae) in the Middle East and Eastern Mediterranean

  • Sepideh Shafaie Department of Botany and Zoology, Faculty of Sciences, Masaryk University, Kotlářská 2, 61137 Brno, Czech Republic
  • Stano Pekár Department of Botany and Zoology, Faculty of Sciences, Masaryk University, Kotlářská 2, 61137 Brno, Czech Republic
  • Osman Seyyar Department of Biology, Faculty of Art & Sciences, Niğde Ömer Halisdemir University, TR-51240 Niğde, Turkey
Keywords: ant-eating spiders, Central Asia, North Africa, new species

Abstract

Although taxonomic research on zodariid spiders has a long history, the identification of many species has remained challenging due to insufficient documentation. In this study, we revisit 50 zodariid species from the Middle East and Eastern Mediterranean. For each taxon examined, we provide detailed photographic documentation of somatic and copulatory structures to facilitate future taxonomic investigations. Seven new species from three different genera are described: Lachesana gavishae Shafaie & Pekár sp. nov. (♂♀, Israel), L. lubinae Shafaie & Pekár sp. nov. (♂♀, Israel), Pax akilae Shafaie & Pekár sp. nov. (♂♀, Syria), Zodarion jakubi Shafaie & Pekár sp. nov. (♂♀, Oman), Z. levyi Shafaie & Pekár sp. nov. (♂♀, Israel and Syria), Z. milani Shafaie & Pekár sp. nov. (♂, Syria), and Z. zonsteini Shafaie & Pekár sp. nov. (♂, Cyprus and Israel). Re-examination of the Z. nicki type specimen confirms its distinction from Z. nitidum, and we provide redescriptions of both sexes of Z. nitidum. Additionally, the male of Z. nicki and the female of Z. namrun are described here for the first time. We revalidate Z. tunetiacum and designate a neotype based on newly examined material. Two new synonymies are also proposed: Acanthinozodium crateriferum syn. nov. is synonymized with A. tibesti, and Z. kunti syn. nov. is synonymized with A. odem. Furthermore, we report ten new country records: A. odem (Syria), A. tibesti (Egypt), Z. abantense (Greece), Z. granulatum (Syria), Z. karpathos (Turkey), Z. morosum (Syria), Z. reticulatum (Turkey), Z. rhodiense (Cyprus, Syria), Z. thoni (Iran, Greece), and Z. tunetiacum (Egypt).

References

Al-Khazali A.M. 2022. Zodarion lutipes (O. Pickard-Cambridge, 1872) (Araneae: Zodariidae) a new record from Iraq. Serket 18 (3): 378–381.

Andreeva E.M. & Tystshenko V.P. 1968. Materials on the fauna of spiders (Aranei) of Tadjikistan. II. Zodariidae. Zoologicheskiĭ Zhurnal 47: 684–689.

Audouin V. 1826. Explication sommaire des planches d’arachnides de l’Égypte et de la Syrie. In: Savigny M.J.C.L. de (ed.) Description de l’Égypte, ou recueil des observations et des recherches qui ont été faites en Égypte pendant l’expédition de l’armée franҫaise, publié par les ordres de sa Majesté l’Empereur Napoléon le Grand. Histoire Naturelle 1 (4): 1–339.

Bosmans R. 1994. Revision of the genus Zodarion Walckenaer, 1833 in the Iberian Peninsula and Balearic Islands (Araneae: Zodariidae). Eos 69: 115–142.

Bosmans R. 1997. Revision of the genus Zodarion Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: Zodariidae). Bulletin of the British Arachnological Society 10 (8): 265–294.

Bosmans R. 2009. Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History 12: 211–295.

Bosmans R. & Gavalas I. 2023. The spiders (Araneae) of the tiny Greek island Iraklia (Kiklades), with the descriptions of 5 new Harpactea species (Araneae: Dysderidae), 3 species new to Europe and 8 new to Greece. Parnassiana Archives 11: 1–91.

Bosmans R., Özkütük R.S., Varli S.V. & Kunt K.B. 2014. Description of a new Zodarion Walckenaer, 1826 from Turkey (Zodariidae; Araneae). Turkish Journal of Zoology 38: 99–101. https://doi.org/10.3906/zoo-1303-11

Bosmans R., Van Keer J., Russell-Smith A., Hadjiconstantis M., Komnenov M., Bosselaers J., Huber S., McCowan D., Snazell R., Decae A., Zoumides C., Kielhorn K.-H. & Oger P. 2019. Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34 (Supplement): 1–173.

Brignoli P.M. 1984. Ragni di Grecia XII. Nuovi dati su varie famiglie (Araneae). Revue suisse de Zoologie 91 (2): 281–321. https://doi.org/10.5962/bhl.part.81881

Caporiacco L. di 1948. L’arachnofauna di Rodi. Redia 33: 27–75.

Chyzer C. & Kulczyński W. 1897. Araneae hungariae. Tomus II. Academia Scientarum Hungaricae, Budapest.

Coşar İ. 2021. A new species of Zodarion (Araneae: Zodariidae) from Turkey. Zoology in the Middle East 67 (3): 259–266. https://doi.org/10.1080/09397140.2021.1957207

Coşar İ. & Danışman T. 2021. Three new Zodarion species (Araneae: Zodariidae) from southeastern Turkey. Zootaxa 5057 (3): 415–428. https://doi.org/10.11646/zootaxa.5057.3.6

Coşar İ. & Danışman T. 2024. A new species of spinibarbe species-group of Zodarion (Araneae: Zodariidae) from Turkey. Arthropoda Selecta 33 (2): 267–272. https://doi.org/10.15298/arthsel.33.2.14

Coşar İ., Danışman T. & Yağmur E.A. 2021. Contributions to the genus Zodarion Walckenaer, 1826 in Turkey, with the description of a new species (Araneae: Zodariidae). Turkish Journal of Zoology 45 (1): 46–53. https://doi.org/10.3906/zoo-2009-36

Coşar İ., Danışman T. & Kunt K.B. 2022. Some new findings of the genus Zodarion (Aranei: Zodariidae) from Turkey. Arthropoda Selecta 31 (2): 228–234. https://doi.org/10.15298/arthsel.31.2.11

Danışman T. & Rubio G.D. 2017. A new species of Zodarion Walckenaer, 1826 (Araneae: Zodariidae) from Turkey. Entomological News 127 (2): 178–183. https://doi.org/10.3157/021.127.0214

Deltshev C.D. 1987. A critical review of genus Zodarion Walckenaer (Araneae, Zodariidae) in Bulgaria. Acta Zoologica Bulgarica 33: 19–25.

Deltshev C., Naumova M., Matevski D. & Indzhov S. 2022. New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species. Zootaxa 5174 (3): 247–261. https://doi.org/10.11646/zootaxa.5174.3.3

Denis J. 1935a. Sur deux araignées de Cyrénaique. Annali del Museo Civico di Storia Naturale di Genova 57: 100–104.

Denis J. 1935b. Les araignées du genre Zodarion Walk., appartenant à la faune d’Italie. Memorie della Società Entomologica Italiana 14: 65–83.

Denis J. 1937. Contribution à l’étude des araignées du genre Zodarion Walckenaer. Festschrift zum 60. Geburstage von Professor Dr. Embrik Strand (Riga) 3: 1–50.

Denis J. 1938. Sur la synonymie de quelques araignées. Bulletin de la Société d’Histoire naturelle de Toulouse 62: 379–389.

Denis J. 1952. Zodariides recuieillis au Maroc et en Mauritanie par M. L. Berland. Bulletin de la Société d’Histoire naturelle d’Afrique du Nord 41: 58–63.

Denis J. 1958. Araignées (Araneidea) de l’Afghanistan. I. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn 120: 81–120.

Denis J. 1962. Araignées recueillies en 1961 par la Mission Belge au Tibesti. Revue de Zoologie et de Botanique africaines 65: 29–32.

Dimitrov D. 1996. A record of Zodarion turcicum Wunderlich from Europe with description of its unknown male (Arachnida, Araneae, Zodariidae). Berichte des naturwissenschaftlich-medizinischen Vereins in Innsbruck 83: 157–158.

Dimitrov D. 2020. Taxonomic contribution to the genus Zodarion Walckenaer, 1826 in Turkey with description of a new species (Araneae: Zodariidae). Zootaxa 4810 (2): 361–367. https://doi.org/10.11646/zootaxa.4810.2.9

Dimitrov D. 2024. On the taxonomy of the Zodarion thoni species-group (Araneae: Zodariidae), with description of a new species with an unmodified embolus. Zootaxa 5419 (1): 112–120. https://doi.org/10.11646/zootaxa.5419.1.4

Drensky P. 1921. Contribution à l’étude des araignées de la Macédoine or et de Pirine Planina. Spisanié na Beulgarskata Akademia na Naoukite 23: 1–80.

Drensky P. 1940. Die Spinnenfauna Bulgariens IV. Mitteilungen aus den königlichen naturwissenschaftlichen Instituten in Sofia 13: 169–194.

Dunin P.M. & Nenilin A.B. 1987. The spider family Zodariidae in the Caucasus (Arachnida: Araneae). Senckenbergiana Biologica 68: 191–198.

Dunlop J.A., Penny D. & Jekel D. 2023. A summary list of fossil spiders and their relatives. In: World Spider Catalog. Version 23.5. Natural History Museum Bern. Available from http://wsc.nmbe.ch [accessed 9 May 2025].

Fage L. 1921. Travaux scientifiques de l’Armee d’Orient (1916–1918). Arachnides. Bulletin du Muséum national d’Histoire naturelle de Paris 27: 96–102, 173–177, 227–232.

Fet V.Y. 1985. [The ecological distribution of spiders in Syunt Khassardagh Reserve]. In: [The Vegetation and Animal Life of Western Kopetdagh]: 271–277. Ashkhabad, Turkmenistan.

Jocqué R. 1987. Descriptions of new genera and species of African Zodariinae with a revision of the genus Heradida (Araneae, Zodariidae). Revue Zoologique africaine 101: 143–163.

Jocqué R. 1991. A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History 201: 1–160.

Jocqué R. 2011. Order Araneae, family Zodariidae. Arthropod fauna of the UAE 4: 15–22.

Jocqué R. & Henrard A. 2015. Revalidation of Acanthinozodium Denis, 1966 with description of three new species and discovery of a remarkable male palpal character (Araneae, Zodariidae). European Journal of Taxonomy 114: 1–23. https://doi.org/10.5852/ejt.2015.114

Jocqué R. & Henrard A. 2020. Three new species of the genus Ranops (Araneae: Zodariidae) from southern Africa. Zootaxa 4899 (1): 186–200. https://doi.org/10.11646/zootaxa.4899.1.10

Karol S. 1969. Eine wenig bekannte Art der Gattung Zodarion (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica 50: 201–203.

Koch C.L. 1872. Die Arachniden Australiens, nach der Natur beschrieben und abgebildet: 105–368. Verlag von Bauer & Raspe, Nürnberg. https://doi.org/10.5962/bhl.title.121660

Kovblyuk M.M. 2003. Spiders of the genus Zodarion (Aranei: Zodariidae) in the fauna of the Crimea. Euroasian Entomological Journal 1 (2): 177–183.

Komnenov M., Pitta E., Zografou K. & Chatzaki M. 2016. Discovering the still unexplored arachnofauna of the National Park of Dadia-Lefkimi-Soufli, NE Greece: a taxonomic review with description of new species. Zootaxa 4096 (1): 1–66. https://doi.org/10.11646/zootaxa.4096.1.1

Krupp F., Khalaf M., Malek M., Streit B. & Al-Jumaily M. 2009. The Middle Eastern biodiversity network: Generating and sharing knowledge for ecosystem management and conservation. Zookeys 31: 3–15. https://doi.org/10.3897/zookeys.31.371

Kulczyński W. 1908. Fragmenta arachnologica. X. Bulletin international de l’Academie des Sciences de Cracovie 1908: 49–86.

Kulczyński W. 1911. Fragmenta Arachnologica. XVI, XVII. Bulletin international de l’Academie des Sciences de Cracovie 1911: 12–75.

Lecigne S. 2011. Inventaire aranéologique dans la Province d’Izmir (Turquie) (Arachnida, Araneae). Le Bulletin d’Arthropoda 46 (2): 5–83.

Lecigne S. 2021. A new species of Sintula (Linyphiidae), redescription of Brigittea innocens (Dictynidae) and eight spider species recorded for Turkey. Arachnologische Mitteilungen 62: 11–34. https://doi.org/10.30963/aramit6204

Lecigne S. 2023. New contribution to the spider fauna (Arachnida: Araneae) of Kerkyra (Corfu) and update of the provisional checklist of species from the Ionian Island. Journal of the Belgian Arachnological Society 38 (1): 1–51.

Lecigne S. & Henrard A. 2022. Zodarion nigrifemur Caporiacco, 1948 newly recorded for the Turkish fauna (Araneae, Zodariidae). Journal of the Belgian Arachnological Society 37 (1): 32–40.

Levy G. 1990. Spiders of the genus Lachesana and a new storenoid genus from Israel (Araneae: Zodariidae). Zoological Journal of the Linnean Society 98 (4): 327–362. https://doi.org/10.1111/j.1096-3642.1990.tb01205.x

Levy G. 1992. The spider genera Palaestina, Trygetus, Zodarion and Ranops (Araneae, Zodariidae) in Israel with annotations on species of the Middle East. Israel Journal of Zoology 38: 67–110.

Levy G. 2007. Calommata (Atypidae) and new spider species (Araneae) from Israel. Zootaxa 1551 (1): 1–30. https://doi.org/10.11646/zootaxa.1551.1.1

Marusik Y.M. & Koponen S. 2001. Spiders of the family Zodariidae from Mongolia (Arachnida: Araneae). Reichenbachia 34: 39–48.

Myers N., Mittermeier R.A., Mittermeier C.G., da Fonseca G.A. & Kent J. 2000. Biodiversity hotspots for conservation priorities. Nature 403: 853–858. https://doi.org/10.1038/35002501

Nae A. 2023. First record for Zodarion morosum (Araneae, Zodariidae), Lessertinella kulczynskii (Araneae, Linyphiidae) and reconfirmation of Palliduphantes milleri (Araneae, Linyphiidae) in the Romanian fauna. Travaux de l’Institut de Spéologie “Émile Racovitza” 62: 55–66. https://doi.org/10.59277/TISER.2023.04

Naumova M., Blagoev G., Dimitrov D., Lazarov S. & Deltshev C. 2017. New data on the spider fauna (Arachnida: Araneae) of Bulgaria. Acta Zoologica Bulgarica 69 (4): 477–481.

Nentwig W., Blick T., Gloor D., Jäger P. & Kropf C. 2020. How to deal with destroyed type material? The case of Embrik Strand (Arachnida: Araneae). Arachnologische Mitteilungen 59 (1): 22–29. https://doi.org/10.30963/aramit5904

Nosek A. 1905. Araneiden, Opilionen und Chernetiden. In: Penther A. & Zederbauer E. (eds) Ergebnisse einer naturwissenschaftlichen Reise zum Erdschias-Dagh (Kleinasien). Annalen des kaiserlich-königlichen Naturhistorischen Hofmuseums in Wien 20 (2–3): 114–154.

Ono H. & Jocqué R. 1986. Two new species of the family Zodariidae from Saudi Arabia (Arachnida: Araneae). Fauna Saudi Arabia 8: 6–11.

Ortiz D., Pekár S., Bilat J. & Alvaraz N. 2021. Poor performance of DNA barcoding and the impact of RAD loci filtering on the species delimitation of an Iberian ant-eating spider. Molecular Phylogenetics and Evolution 154: e106997. https://doi.org/10.1016/j.ympev.2020.106997

Ortiz D., Pekár S. & Dianat M. 2024. A consequential one-night stand: Episodic historical hybridization leads to mitochondrial takeover in sympatric desert ant-eating spiders. Molecular Phylogenetics and Evolution 199: e108167. https://doi.org/10.1016/j.ympev.2024.108167

Ovtchinnikov S.V. 2006. New genus and species of spiders of the subfamily Zodariinae (Araneae, Zodariidae) from Pakistan. Vestnik Zoologii 40: 77–79.

Ovtchinnikov S.V., Ahmad B. & Gurko V.O. 2009. Parazodarion, a new genus of the spider family Zodariidae (Araneae) from Asia. Vestnik Zoologii 43: 471474.

Özkütük R.S. 2022. A new ant-eating spider record (Aranei: Zodariidae) from Turkey. Caucasian Entomological Bulletin 18 (2): 239–242. https://doi.org/10.23885/181433262022182-239242

Pekár S., Šmerda J., Hrušková M., Šedo O., Muster C., Cardoso P., Zdráhal Z., Korenko S., Bureš P., Líznarová E. & Sentenská L. 2012. Prey-race drives differentiation of biotypes in ant-eating spiders. Journal of Animal Ecology 81 (4): 838–848. https://doi.org/10.1111/j.1365-2656.2012.01957.x

Pekár S., Cardoso P. & Meierrose C. 2003. Additions to the knowledge of Portuguese zodariid spiders (Araneae: Zodariidae). Bulletin of the British Arachnological Society 12 (9): 385–395.

Pekár S., Ortiz D., Sentenská L. & Šedo O. 2022. Ecological specialization and reproductive isolation among closely related sympatric ant-eating spiders. Journal of Animal Ecology 91 (9): 1855–1868. https://doi.org/10.1111/1365-2656.13767

Pickard-Cambridge F. 1902. A revision of the genera of Araneae or spiders with reference to their type species. Annals and Magazine of Natural History 7 (9): 5–20. https://doi.org/10.1080/00222930208678530

Pickard-Cambridge O. 1872. General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London 40 (1): 212–354. Available from https://ia801209.us.archive.org/34/items/biostor-99678/biostor-99678.pdf [accessed 18 Jul. 2025].

Pickard-Cambridge O. 1876. Catalogue of a collection of spiders made in Egypt, with descriptions of new species and characters of a new genus. Proceedings of the Zoological Society of London 44 (3): 541–630. https://doi.org/10.1111/j.1096-3642.1876.tb02595.x

Ramos M., Coddington J.A., Christenson T.E. & Irschick D.J. 2005. Have male and female genitalia coevolved? A phylogenetic analysis of genitalic morphology and sexual size dimorphism in web-building spiders (Araneae: Araneoidea). Evolution 59 (9):1989–1999. https://doi.org/10.1554/04-499.1

Roewer C.F. 1928. Araneae. In: Zoologische Streifzüge in Attika, Morea, und besonders auf der Insel Kreta, II. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 27: 92–123.

Roewer C.F. 1960. Solifugen und Opilioniden - Araneae Orthognathae, Haplogynae und Entelegynae (Contribution à l’étude de la faune d’Afghanistan 23). Göteborgs Kungliga Vetenskaps och Vitterhets Samhälles Handlingar 8 (7): 1–53.

Seropian A., Bulbulashvili N., Otto S., Krammer H.-J., Kachlishvili N. & Datunashvili A. 2023. Picking pearls from the Silk Road: insights into the spider (Arachnida, Araneae) diversity in Georgia from the Caucasus barcode of life project. Part II. Caucasiana 2: 231–297. https://doi.org/10.3897/caucasiana.2.e110536

Seyyar O., El-Hennawy H.K., Demir H. & Türkeş T. 2008. The first record of genus Pax (Araneae: Zodariidae) in Turkey. Serket 11: 51–54.

Shafaie S., Pekár S. & Ortiz D. 2025. Integrative taxonomy of the Iberian Zodarion species of the rubidum and styliferum groups (Araneae: Zodariidae). Zootaxa 5624 (1): 1–69. https://doi.org/10.11646/zootaxa.5624.1.1

Simon E. 1864. Histoire naturelle des Araignées (Aranéides). Roret, Paris. https://doi.org/10.5962/bhl.title.47654

Simon E. 1870. Sur les aranéides de la famille des Enydes qui habitant l’Espagne et le Maroc. Revue et Magasin de Zoologie pure appliqée 2 (22): 51–54, 97–103, 142–148.

Simon E. 1873. Aranéides nouveaux ou peu connus du midi de l’Europe. Extrait des Mémoires de la Société royale des Sciences de Liège 2 (5): 187–351. [separately paginated version (pp. 1–174) seen and cited here.] https://doi.org/10.5962/bhl.title.124166

Simon E. 1885. Études sur les Arachnides recueillis en Tunisie en 1883 et 1884 par MM. A. Letourneux, M. Sédillot et Valéry Mayet, membres de la mission de l’Exploration scientifique de la Tunisie. In: Exploration scientifique de la Tunisie, publiée sous les auspices du Ministère de l’instruction publique. Zoologie – Arachnides. Paris.

Simon E. 1889. Arachnidae transcaspicae ab ill. Dr. G. Radde, Dr. A. Walter et A. Conchin inventae (annis 1886–1887). Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 39: 373–386.

Simon E. 1890. Études arachnologiques. 22e Mémoire. XXXIV. Étude sur les arachnides de l’Yemen. Annales de la Société entomologique de France 6 (10): 77–124.

Simon E. 1893. Histoire naturelle des Araignées. Deuxième édition, tome premier. Roret, Paris. https://doi.org/10.5962/bhl.title.51973

Simon E. 1908. Étude sur les arachnides recueillis par M. le Dr. Klaptocz en Tripolitaine. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere 26: 419–438.

Stefanovska D., Naumova M., Prelik D., Deltshev C. & Lazarov S. 2008. Spiders from the Skopje region: a faunistic and zoogeographical analysis. Historia Naturalis Bulgarica 19: 35–49.

Strand E. 1906. Diagnosen nordafrikanischer, hauptsächlich von Carlo Freiherr von Erlanger gesammelter Spinnen. Zoologischer Anzeiger 30: 604–637, 655–690.

Strand E. 1908. Nordafrikanische Spinnen, hauptsächlich von Carlo Freiherr von Erlanger gesammelt (Dictynidae, Eresidae, Sicariidae, Dysderidae, Caponiidae, Palpimanidae, Zodariidae, Urocteidae, Pholcidae, Agelenidae, Pisauridae). Archiv für Naturgeschichte 74: 67–128. https://doi.org/10.5962/bhl.part.37388

Strand E. 1914. Zweite Mitteilung über Spinnen aus Palästina, gesammelt von Herrn Dr J. Aharoni. Archiv für Naturgeschichte 80 (A3): 173–186. https://doi.org/10.5962/bhl.part.26478

Thorell T. 1869. On European spiders. Part I. Review of the European genera of spiders, preceded by some observations on zoological nomenclature. Nova Acta Regiae Societatis Scientiarum Upsaliensis 7: 1–108.

Thorell T. 1873. Remarks on Synonyms of European Spiders. Part IV: 375–645. C.L. Lundström, Uppsala. https://doi.org/10.5962/bhl.title.69282

Thorell T. 1881. Studi sui Ragni Malesi e Papuani. III. Ragni dell’Austro Malesia e del Capo York, conservati nel Museo civico di storia naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova 17: 1–720.

Vlassov J.P. & Sytshevskaja V.I. 1937. The spiders of the neighbourhood of Ashkhabad. Trudy Soveta po Izuceniju Proizvoditeljnych Sil, Serija Turkmenskaja 9: 247–258.

Walckenaer C.A. 1826. Aranéides. In: Faune française ou histoire naturelle générale et particulière des animaux qui se trouvent en France, constamment ou passagèrement, à la surface du sol, dans les eaux qui le baignent et dans le littoral des mers qui le bornent par Viellot, Desmarrey, Ducrotoy, Audinet, Lepelletier et Walckenaer 11–12: 1–96. Livraisou, Paris.

Walckenaer C.A. 1837. Histoire naturelle des Insectes. Aptères. Tome premier. Roret, Paris. https://doi.org/10.5962/bhl.title.61095

Walckenaer C.A. 1841. Histoire naturelle des Insectes. Aptères. Tome deuxième. Roret, Paris. https://doi.org/10.5962/bhl.title.61095

Walckenaer C.A. 1847. Dernier Supplément. In: Walckenaer C.A. & Gervais P. (eds) Histoire naturelle des Insectes. Aptères. Tome quatrième: 365–596. Roret, Paris. https://doi.org/10.5962/bhl.title.61095

Weiss I. 1987. Zodarion geticum n. sp., eine Spinne mit Duftorganen aus Rumänien (Arachnida, Araneae, Zodariidae). Reichenbachia 25: 103–106.

World Spider Catalog 2025. World Spider Catalog. Version 26. Natural History Museum Bern. Available from http://wsc.nmbe.ch [accessed 9 May 2025]. https://doi.org/10.24436/2

Wunderlich J. 1973. Beschreibung einiger bisher unbekannter Arten der Gattung Zodarion Walckenaer aus Südeuropa (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica 54: 171–176.

Wunderlich J. 1980. Zur Gattung Zodarion Walckenaer 1847 mit Neubeschreibungen (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica 60 (3/4): 229–240.

Wunderlich J. 2022. Some spiders of (Araneae) of the western Palearctic. Beiträge zur Araneologie 15: 4–78, 199.

Zamani A. & Marusik Y.M. 2021. Revision of the spider family Zodariidae (Arachnida, Araneae) in Iran and Turkmenistan, with seventeen new species. ZooKeys 1035: 145–193. https://doi.org/10.3897/zookeys.1035.65767

Zamani A. & Marusik Y.M. 2022. New taxonomic considerations in Zodariellum Andreeva & Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera. Zootaxa 5178 (2): 161–177. https://doi.org/10.11646/zootaxa.5178.2.3

Zonstein S.L. 2025. A redescription of Acanthinozodium odem (Levy, 2007) comb. nov. (Araneae, Zodariidae), with first description of the male. Zootaxa 5583 (2): 391–397. https://doi.org/10.11646/zootaxa.5583.2.10

Zonstein S.L. & Ovtchinnikov S.V. 1999. A new Central Asian species of the spider genus Lachesana Strand, 1932 (Araneae, Zodariidae: Lachesaninae). Tethys Entomological Research 1: 59–62.

Published
2025-09-18
How to Cite
Shafaie, S., Pekár, S., & Seyyar, O. (2025). New data on zodariid spiders (Arachnida, Araneae) in the Middle East and Eastern Mediterranean. European Journal of Taxonomy, 1016(1), 1-139. https://doi.org/10.5852/ejt.2025.1016.3063
Section
Monograph