New annulate Carboniferous orthoconic and cyrtoconic cephalopods of the family Brachycycloceratidae Furnish, Glenister & Hartman, 1962

Keywords: Orthoceratoidea, Brachycycloceratidae, Carboniferous, Rhenish Mountains

Abstract

The Carboniferous orthoconic or slightly cyrtoconic cephalopod family Brachycycloceratidae Furnish, Glenister & Hansman, 1962 is well described based on material from North America, whereas the Early Carboniferous European occurrences of this family are much less well-known. Almost all records from the Rhenish Mountains have been assigned to the single species Brachycycloceras scalare (d’Archiac & de Verneuil, 1842). This species is revised here and used as the type species of the genus Rhenocycloceras gen. nov. The new Early Carboniferous species Rhenocycloceras conicum sp. nov. and R. denckmanni sp. nov. from the Rhenish Mountains of Germany, R. macdiarmadai sp. nov. from Co. Leitrim in Ireland and R. africanum sp. nov. from the Anti-Atlas of Morocco are described. Additionally, two new Early Carboniferous species of Caneycycloceras Niko & Mapes, 2011 are described from the Rhenish Mountains; these are C. rotersorum sp. nov. and C. fuerstenbergorum sp. nov. The early Late Carboniferous species B. koninckianum (d’Orbigny, 1849) is discussed on the basis of material from the type locality in Belgium.

References

Aubrechtová M., Turek V. & Evans D. 2025. New reticulated and annulated cephalopods from the Ordovician of the Prague Basin (Central Bohemia). Journal of Paleontology 99: 890–904. https://doi.org/10.1017/jpa.2025.10142

Bednarczyk W., Korejwo K., Łobanowski H. & Teller L. 1968. Stratygrafia utworów paleozoicznych z wiercenia Strożyska 5 (niecka miechowska). Acta Geologica Polonica 18: 677–690.

Brandon A. & Hodson F. 1984. The stratigraphy and palaeontology of the Late Visean and Early Namurian rocks of north-east Connought. The Geological Survey of Ireland, Special Paper 6: 1–54.

Brown T. 1841. Description of some new species of fossil shells, found chiefly in the Vale of Todmorden, Yorkshire. Transactions of the Manchester Geological Society 1: 211–232.

Cichowolski M. 2008. The orthocerid Dawsonoceras? (Nautiloidea) from the Lipeón Formation (Silurian, Northwestern Argentina). Ameghiniana 45: 791–793.

Cramer L.W. 1827. Geognostische Fragmente von Dillenburg und der umliegenden Gegend. Heyer, Giessen.

d’Archiac V. & de Verneuil M.E. 1842. On the fossils of the older deposits in the Rhenish Provinces; preceded by a general survey of the fauna of the Palaeozoic Rocks, and followed by a tabular list of the organic remains of the Devonian. Transactions of the Geological Society of London 6: 303–410. https://doi.org/10.1144/transgslb.6.2.303

d’Orbigny A. 1849. Prodrôme de paléontologie stratigraphique universelle des animaux mollusques et rayonnés au cours élémentaire de paléontologie et de géologie stratigraphiques. Premier volume. Masson, Paris. https://doi.org/10.5962/bhl.title.45605

de Koninck L.G. 1844. Description des animaux fossiles qui se trouvent dans le terrain carbonifère de la Belgique. H. Dessain, Brussels. Available from https://www.biodiversitylibrary.org/item/317806 [accessed 27 Jan. 2026].

de Koninck L.G. 1880. Faune du calcaire carbonifère de la Belgique. Deuxième partie: Genres Gyroceras, Cyrtoceras, Gomphoceras, Orthoceras, Subclymenia et Goniatites. Annales du Musée royal d’Histoire naturelle de Belgique 5: 1–333. Available from https://www.biodiversitylibrary.org/item/250929 [accessed 27 Jan. 2026].

Demanet F. 1941. Faune et stratigraphie de l’étage namurien de la Belgique. Mémoires du Musée royal d’Histoire naturelle de Belgique 97: 1–327.

Dernov V. 2021. New finds of the Carboniferous cephalopods on the territory of Ukraine. Visnyk of VN Karazin Kharkiv National University, series Geology. Geography. Ecology 55: 72–81. https://doi.org/10.26565/2410-7360-2021-55-05

Dernov V. 2024. A low-diverse Visean (Mississippian) cephalopod assemblage from the Dnipro-Donets Depression (Ukraine). Revista Brasileira de Paleontologia 27: e20240433. https://doi.org/10.4072/rbp.2024.2.0433

Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica 45: 1–219.

Elias M.K. 1958. Late Mississippian Fauna from the Redoak Hollow Formation of Southern Oklahoma: Pt. 4: Gastropoda, Scaphopoda, Cephalopoda, Ostracoda, Thoracica, and Problematica. Journal of Paleontology 32: 1–57.

Evans D.H. 1994. The cephalopod fauna of the Bardahessaigh Formation (Caradoc Series) of Pomeroy, County Tyrone. Irish Journal of Earth Sciences 13: 11–29.

Fleming J. 1815. Observations on the Orthoceratites of Scotland. Annals of Philosophy 5: 199–206.

Fleming J. 1828. A History of British Animals, Exhibiting the Descriptive Characters and Systematical Arrangement of the Genera and Species of Quadrupeds, Birds, Reptiles, Fishes, Mollusca, and Radiata of the United Kingdom: Including the Indigenous, Extirpated, and Extinct Kinds, together with Periodical and Occasional Visitants. Bell, Bradfute, Duncan, Edinburgh, London. https://doi.org/10.5962/bhl.title.12859

Flower R.H. 1964. Nautiloid shell morphology. Memoirs of the State Bureau of Mines and Mineral Resources, New Mexico Institute of Mining and Technology 13: 1–79. https://doi.org/10.58799/M-13

Foerste A.F. 1928. American Arctic and related cephalopods. Bulletin of the Scientific Laboratories of Denison University 23: 1–110.

Foerste A.F. 1932. Black River and other cephalopods from Minnesota, Wisconsin, Michigan, and Ontario (Part 1). Bulletin of the Scientific Laboratories of Denison University 27: 47–136.

Foord A.H. 1888. Catalogue of the Fossil Cephalopoda in the British Museum. Part 1: Containing Part of the Suborder Nautiloidea, Consisting of the Families Orthoceratidæ, Endoceratidæ, Actinoceratidæ, Gonioceratidæ, Ascoceratidæ, Poterioceratidæ, Cyrtoceratidæ, and Supplement. Order of the Trustees, London. Available from https://www.biodiversitylibrary.org/item/196423 [accessed 27 Jan. 2026].

Foord A.H. 1897. Monograph on the Carboniferous Cephalopoda of Ireland. Part I. Family Orthoceratidæ (in part). Monographs of the Palæontographical Society 51: 1–22. https://doi.org/10.1080/02693445.1897.12028044

Fuchs A. 1915. Der Hunsrückschiefer und die Unterkoblenzschichten am Mittelrhein (Loreleigegend). Abhandlungen der Königlich Preußischen geologischen Landesanstalt, Neue Folge 79: 1–81.

Furnish W.M., Glenister B.F. & Hansman R.H. 1962. Brachycycloceratidae, novum, deciduous Pennsylvanian nautiloids. Journal of Paleontology 36: 1341–1356.

Girty G.H. 1909. The fauna of the Caney Shale of Oklahoma. Bulletin of the U.S. Geological Survey 377: 1–106.

Girty G.H. 1911. On some new genera and species of Pennsylvanian fossils from the Wewoka formation of Oklahoma. New York Academy of Science, Annals 2: 119–156. https://doi.org/10.1111/j.1749-6632.1911.tb56931.x

Girty G.H. 1915. Fauna of the Wewoka Formation of Oklahoma. Bulletin of the U.S. Geological Survey 544: 1–353.

Gordon M. 1965. Carboniferous cephalopods of Arkansas. Professional Papers, U.S. Geological Survey 460: 1–322. https://doi.org/10.3133/pp460

Gunia T. & Górecka T. 1960. O nowym stanowisku fauny dolnokarbońskiej w okolicy Wojborza na Dolnym Śląsku. Rocznik Polskiego Towarzystwa Geologicznego 30: 315–326.

Hind W. 1905. Notes on the homotaxial equivalents of the beds which immediately succeed the Carboniferous Limestone in the west of Ireland. Proceedings of the Royal Irish Academy, B 25: 93–116.

Histon K. 1991. Cycloceras M‘Coy, 1844 (Mollusca, Nautiloidea): proposed designation of C. laevigatum M‘Coy, 1844 as the type species, and proposed designation of a neotype for C. laevigatum. Bulletin of Zoological Nomenclature 48: 97–99. https://doi.org/10.5962/bhl.part.691

Histon K. 1997. Nautiloid species from the Carboniferous of Ireland. Irish Journal of Earth Sciences 16: 45–60.

Holzapfel E. 1889. Die Cephalopoden-führenden Kalke des unteren Carbon von Erdbach-Breitscheid bei Herborn. Palaeontologische Abhandlungen, Neue Folge 5: 1–74.

Horný R. 1956. On the genus Dawsonoceras Hyatt, 1884 (Nautiloidea) of Central Bohemia. Sborník Ústředního Ústavu geologického, Oddíl paleontologický 22: 425–452.

Hüffner E. 1914. Beiträge zur Kenntnis des deutschen Culms. Jahrbuch der Königlich Preußischen Geologischen Landesanstalt 35: 448–548.

ICZN 1993. Cycloceras M‘Coy, 1844 (Mollusca, Nautiloidea): C. laevigatum M‘Coy, 1844 designated as the type species, and a neotype designated for C. laevigatum. Bulletin of Zoological Nomenclature 50: 159–160.

King A.H. & Evans D.H. 2019. High-level classification of the nautiloid cephalopods: a proposal for the revision of the Treatise Part K. Swiss Journal of Palaeontology 138: 65–85. https://doi.org/10.1007/s13358-019-00186-4

Klug C., Döring S., Korn D. & Ebbighausen V. 2006. The Visean sedimentary succession at the Gara el Itima (Anti-Atlas, Morocco) and its ammonoid faunas. Fossil Record 9: 3–60. https://doi.org/10.5194/fr-9-3-2006

Klug C., Korn D., Landman N.H., Tanabe K., De Baets K. & Naglik C. 2015. Describing ammonoid conchs. In: Klug C., Korn D., De Baets K., Kruta I. & Mapes R.H. (eds) Ammonoid Paleobiology: From macroevolution to paleogeography, Topics in Geobiology 44: 3–24. Springer, Dordrecht. https://doi.org/10.1007/978-94-017-9630-9_1

Korn D. 1988. Die Goniatiten des Kulmplattenkalkes (Cephalopoda, Ammonoidea; Unterkarbon; Rheinisches Schiefergebirge). Geologie und Paläontologie in Westfalen 11: 1–293.

Korn D. 1996. Revision of the Rhenish Late Viséan goniatite stratigraphy. Annales de la Société géologique de Belgique 117: 129–136.

Kröger B. & Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 137–163. https://doi.org/10.1002/mmng.200600005

Kuhn O. 1940. Paläozoologie in Tabellen. Fischer, Jena.

M‘Coy F. 1844. A Synopsis of the Characters of the Carboniferous Limestone Fossils of Ireland. University Press, Dublin. https://doi.org/10.5962/bhl.title.11559

Manda Š. & Turek V. 2015. Colour patterns on Silurian orthocerid and pseudorthocerid conchs from Gotland palaeoecological implications. Estonian Journal of Earth Sciences 64: 74–79. https://doi.org/10.3176/earth.2015.13

Meek F.B. & Worthen A.H. 1860. Descriptions of new Carboniferous fossils from Illinois and other western States. Proceedings of the Academy of Natural Sciences Philadelphia 12: 447–472.

Meek F.B. & Worthen A.H. 1862. Descriptions of new Palæozoic Fossils from Illinois and Iowa. Proceedings of the Academy of Natural Sciences Philadelphia 13: 128–148.

Miller A.K. & Owen J.B. 1934. Cherokee nautiloids of the northern Mid-Continent region. University of Iowa Studies in Natural History 16: 185–272.

Miller A.K. & Unklesbay A.G. 1942. The cephalopod fauna of the Conemaugh Series in Western Pennsylvania. Annals of the Carnegie Museum 29: 127–174. https://doi.org/10.5962/p.215157

Miller A.K., Dunbar C. & Condra G.E. 1933. The nautiloid cephalopods of the Pennsylvanian system in the Mid-Continent region. Nebraska Geological Survey Bulletin 9: 1–240.

Münster G. Graf z. 1840. Die Versteinerungen des Uebergangskalkes mit Clymenien und Orthoceratiten von Oberfranken. Beiträge zur Petrefactenkunde 3: 33–121.

Nebe B. 1911. Die Culmfauna von Hagen i. W., ein Beitrag zur Kenntnis des westfälischen Untercarbons. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Beilage-Band 31: 421–495.

Niko S. & Ehiro M. 2025. Changhsingian (latest Permian) orthoconic cephalopods from the South Kitakami Belt, Northeast Japan. Bulletin of the Tohoku University Museum 24: 1–4.

Niko S. & Mapes R.H. 2009. Redescription and new information on the Carboniferous cephalopod Brachycycloceras normale Miller, Dunbar and Condra, 1933. Paleontological Research 13: 337–343. https://doi.org/10.2517/1342-8144-13.4.337

Niko S. & Mapes R.H. 2011. Caneycycloceras, a new brachycycloceratid cephalopod genus from the Early Carboniferous in Oklahoma, Midcontinent North America. Paleontological Research 15: 292–295. https://doi.org/10.2517/1342-8144-15.4.292

Niko S., Nishida T. & Kyama Y. 1987. Middle Carboniferous Orthocerataceae and Pseudorthocerataceae (Mollusca: Cephalopoda) from the Akiyoshi Limestone, Yamaguchi Prefecture (Molluscan Paleontology of the Akiyoshi Limestone Group-VIII). Transactions and Proceedings of the Paleontological Society of Japan. New series 1987: 335–345.

Niko S., Pillevuit A. & Nishida T. 1996. Early Late Permian (Wordian) non-ammonoid cephalopods from the Hamrat Duru Group, central Oman Mountains. Transactions and Proceedings of the Paleontological Society of Japan. New series 1996: 522–527.

Patteisky K. 1929. Die Geologie und Fossilführung der mährisch-schlesischen Dachschiefer- und Grauwackenformation. Naturwissenschaftlicher Verein in Troppau, Opava.

Phillips D. 1985. The nautiloid Brachycycloceras in the Upper Carboniferous of Britain. Palaeontology 28: 235–242.

Phillips J. 1836. Illustrations of the Geology of Yorkshire. Part 2: The Mountain Limestone District. John Murray, London.

Pohle A., Kröger B., Warnock R., King A.H., Evans D.H., Aubrechtová M., Cichowolski M., Fang X. & Klug C. 2022. Early cephalopod evolution clarified through Bayesian phylogenetic inference. BMC biology 20: 1–30. https://doi.org/10.1186/s12915-022-01284-5

Roemer F. 1870. Geologie von Oberschlesien. Nischkowsky.

Roemer F. 1876. Lethaea palaeozoica oder Beschreibung und Abbildung der für die Gebirgs-Formationen bezeichnendsten Versteinerungen. I. Theil. Lethaea palaeozoica. Schweizerbart, Stuttgart.

Roemer F.A. 1852. Beiträge zur geologischen Kenntniss des nordwestlichen Harzgebirges. Zweite Abtheilung. Palaeontographica 3: 69–111.

Sandberger G. & Sandberger F. 1850–1856. Die Versteinerungen des rheinischen Schichtensystems in Nassau. Mit einer kurzgefassten Geognosie dieses Gebietes und mit steter Berücksichtigung analoger Schichten anderer Länder. Kreidel & Niedner, Wiesbaden. https://doi.org/10.5962/bhl.title.52349

Sarres J.H. 1857. De petrefactis, quae in schisto posidonico prope Elberfeldam urbem inveniuntur: dissertatio inauguralis. Friedrich-Wilhelm-Universität, Berlin. https://doi.org/10.5962/bhl.title.132569

Schmidt H. 1956. Orthocone Cephalopoden aus dem deutschen Unterkarbon. Paläontologische Zeitschrift 30: 41–68. https://doi.org/10.1007/BF03041872

Shimansky V.N. 1965. Nautiloidei. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 40–47.

Shimansky V.N. 1968. Kamennougol'nye Orthoceratida, Oncoceratida, Actinoceratida i Bactritida. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 117: 1–151.

Sommer K. 1910. Die Fauna des Culm von Königsberg bei Gießen. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Beilage-Band 28: 611–660.

Sowerby J. 1815–1818. The Mineral Conchology of Great Britain: or Coloured Figures and Descriptions of those Remains of Testaceous Animals or Shells, which have been Preserved at Various Times and Depths in the Earth. Vol. 2. Meredith, London. Available from https://www.biodiversitylibrary.org/item/210427 [accessed 27 Jan. 2026].

Sweet W.C. 1964. Cephalopoda–General Features. In: Moore R.C. (ed.) Treatise on Invertebrate Paleontology: K4–K13. The Geological Society of America and The University of Kansas Press, Lawrence.

Teichert C. 1964. Morphology of hard parts. In: Moore R.C. (ed.) Treatise on Invertebrate Paleontology: K13–K53. The Geological Society of America and The University of Kansas Press, Lawrence.

Teichert C. 1967. Major Features of Cephalopod Evolution. In: Teichert C. & Yochelson E.-L. (eds) Essays in Paleontology and Stratigraphy, Special Publication: 162–210. The University of Kansas Press, Lawrence.

Turek V. & Aubrechtová M. 2024. Micro-CT reveals 3D endosiphuncular structure in Late Ordovician actinoceratid cephalopod from the Prague Basin (Czech Republic). Bulletin of Geosciences 99 (3): 169–189. https://doi.org/10.3140/bull.geosci.1901

von Koenen A. 1879. Die Kulm-Fauna von Herborn. Neues Jahrbuch für Mineralogie, Geologie und Palaeontologie 1879: 309–346.

von Meyer H. 1831. Beschreibung des Orthoceratites striolatus und über den Bau das Vorkommen einiger vielkammerigen fossilen Cephalopoden; nebst der Beschreibung von Calymene aequalis. Verhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Akademie der Naturforscher 15: 59–113.

Wang Q., Korn D., Nemyrovska T. & Qi Y. 2018. The Wenne river bank section–an excellent section for the Viséan–Serpukhovian boundary based on conodonts and ammonoids (Mississippian; Rhenish Mountains, Germany). Newsletters on Stratigraphy 51: 427–444. https://doi.org/10.1127/nos/2018/0440

Worthen A.H. 1882. Corrections and proposed new names for species described in the Geological Reports of Illinois under names that were preoccupied; and descriptions of two new species of fossil shells from the measures of Illinois and Kansas. Bulletin of the Illinois State Museum of Natural History 1: 38–40.

Żakowa H. 1958. Biostratygrafia utworów morskich Dolnego Karbonu z obszaru Walbrzycha Miasta na Dolnym Slasko. Prace Instytutu Geologicznego 19: 1–211.

Żakowa H. 1960. Nowe stanowisko fauny górnowizenskiej z Walbrzycha Miasta. Kwartalnik geologiczny 4: 619–630.

Published
2026-03-25
How to Cite
Korn, D., & Aubrechtová, M. (2026). New annulate Carboniferous orthoconic and cyrtoconic cephalopods of the family Brachycycloceratidae Furnish, Glenister & Hartman, 1962. European Journal of Taxonomy, 1048(1), 1-39. https://doi.org/10.5852/ejt.2026.1048.3231
Section
Research article