A new genus of mongoliulid millipedes from the Far East of Russia, with a list of species in the family (Diplopoda, Julida, Mongoliulidae)

Henrik Enghoff, Laura Mark Jensen, Elena V. Mikhaljova


The genus Koiulus gen. nov. and its type-species, Koiulus interruptus gen. et sp. nov., are described from the Russian Far East. The new genus is compared with other genera of Mongoliulidae, in particular with Ussuriiulus Golovatch, 1980, also from the Russian Far East, with which it shares the absence of ozopores from individual body rings distributed along the body, a condition so far otherwise unknown in the superorder Juliformia. A synoptic table of genera and a list of species of Mongoliulidae are presented.


taxonomy; new species

Full Text:


DOI: https://doi.org/10.5852/ejt.2017.326


Attems C. 1901. Myriopoden. In: Horváth G. (ed.) Zoologische Ergebnisse der dritten asiatischen Forschungsreise des Grafen Zichy 2: 275–310. Hornyánszky, Budapest; Hiersemann, Leipzig.

Attems C. 1909. Die Myriopoden der Vega-Expedition. Arkiv för Zoologi 5 (3): 1–84.

Enghoff H. 1981. A cladistic analysis and classification of the millipede order Julida. Zeitschrift für zoologische Systematik und Evolutionsforschung 19: 285–319.

Enghoff H. 1982. The millipede genus Cylindroiulus on Madeira – an insular species swarm (Diplopoda, Julida: Julidae). Entomologica Scandinavica, Supplement 18: 1–142.

Enghoff H., Petersen G. & Seberg O. 2011. Phylogenetic relationships in the millipede family Julidae. Cladistics 27: 606–616. https://doi.org/10.1111/j.1096-0031.2011.00360.x

Enghoff H., Petersen G. & Seberg O. 2013. The aberrant millipede genus Pteridoiulus and its position in a revised molecular phylogeny of the family Julidae (Diplopoda, Julida). Invertebrate Systematics 27: 515–529. https://doi.org/10.1071/IS13016

Enghoff H., Golovatch S., Short M., Stoev P. & Wesener T. 2015. Diplopoda – Taxonomic overview. In: Minelli A. (ed.) The Myriapoda 2. Treatise on Zoology – Anatomy, Taxonomy, Biology: 363–453. Brill, Leiden, Boston.

Golovatch S.I. 1979a. A new genus of the superfamily Parajuloidea (Diplopoda) new to the USSR. Entomologicheskoe Obozrenie 58 (4): 904–908 [in Russian, English summary].

Golovatch S.I. 1979b. Three new to the USSR fauna genera of Diplopoda-Chilognatha. Zoologicheski Zhurnal 58 (3): 336–343 [in Russian, English summary].

Golovatch S.I. 1980a. A contribution to the millipede fauna of Korea (Diplopoda). Folia Entomologica Hungarica 41 (1): 49–58.

Golovatch S.I. 1980b. New forms of Diplopoda from the Soviet Far East and their zoogeographical relationships. Zoologicheski Zhurnal 59 (2): 199–207 [in Russian, English summary].

Hoffman R.L. & Howell K.M. 1983. Dendrostreptus, a new genus for an arboreal Tanzanian milliped, with notes on related forms. Revue de Zoologie africaine 97 (3): 625–632.

Hoffman R.L. & Howell K.M. 1995. On the status of Microtrullius Attems, 1950, an enigmatic genus of the diplopod family Spirostreptidae. Journal of African Zoology 109: 173–184.

Hoffman R.L. & Howell K.M. 1996. Synopsis of Macrolenostreptini, a new tribe of Tanzanian spirostreptid millipeds (Diplopoda Spirostreptida). Tropical Zoology 9: 441–453. https://doi.org/10.1080/03946975.1996.10539322

Lim K.Y. & Mikhaljova E.V. 2001. A new species of the millipede genus Skleroprotopus Attems, 1901 (Diplopoda: Julida: Mongoliulidae) from Korea. Arthropoda Selecta 9 (2): 119–122.

Mikhaljova E.V. 1997. Review of the cavernicolous millipede fauna of the Far East of Russia, with description of a new troglophilic species (Diplopoda). Arthropoda Selecta 5 (3–4): 143–149.

Mikhaljova E.V. 1998. The millipedes of the Far East of Russia (Diplopoda). Arthropoda Selecta 7 (1): 1–77.

Mikhaljova E.V. & Korsós Z. 2003. Millipedes (Diplopoda) from Korea, the Russian Far East, and China in the collection of the Hungarian Natural History Museum. Acta Zoologica Scientiarum Hungaricae 49 (3): 215–242.

Mikhaljova E.V. & Lim K.Y. 2001. New millipedes from Korea, with notes on identity of Epanerchodus koreanus Verhoeff, 1937 (Diplopoda). Arthropoda Selecta 10 (1): 19–26.

Miyosi Y. 1957. Beiträge zur Kenntnis japanischer Myriopoden. 19. Aufsatz: Über eine neue Gattung und zwei neue Arten von Diplopoden. Dobutsugako Zasshi [The Zoological Magazine] 66: 29–33 [in Japanese, German summary].

Murakami Y. 1971. The fauna of the insular lava caves in West Japan. X. Myriapoda. Bulletin of the National Science Museum, Tokyo 14 (3): 311–332.

Murakami Y. 1990. The millipeds of the genus Kopidoiulus (Diplopoda, Julida, Mongoliulidae). Journal of the Speleological Society of Japan 15: 1–14.

Shinohara K. 1960. Three new species of Juloidea (Diplopoda) from Chichibu. Bulletin of the Chichibu Museum of Natural History 10: 23–30 [in Japanese, English summary].

Takakuwa Y. 1940a. Über eine neue Diplopodengattung aus Japan. Transactions of the Sapporo Natural History Society 16 (3): 117–119.

Takakuwa Y. 1940b. Über Scleroprotopus-Arten (Chilopoda) [sic]. Annotationes Zoologicae Japonenses 19 (1): 19–22.

Takakuwa Y. 1941. Eine neue Gattung der Paraiulidae (Diplopoda). Zoological Magazine, Tokyo 53: 364–366.

Takakuwa Y. 1942. Zur Kenntnis der japanischen Diplopoden. Annotationes Zoologicae Japonenses 21 (1): 39–47.

Takakuwa Y. 1954. Diplopoden aus Japan und ihm angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo. [in Japanese].

Takakuwa Y. & Takashima H. 1949. Myriapods collected in Shansi, North China. Acta Arachnologica 11 (3–4): 51–69 [in Japanese, English summary].

Zhang C.Z. 1985. A new millipede of the genus Skleroprotopus in Stone Buddha Cave, Fangsheng County, Beijing. In: Karst Geomorphology and Speleology: 154–156. Science Press, Beijing [in Chinese].


  • There are currently no refbacks.

License URL: https://creativecommons.org/licenses/by/3.0/