The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France)
Abstract
References
Bourgoin T. 2020. FLOW (Fulgoromorpha Lists On the Web): a World Knowledge Base dedicated to Fulgoromorpha. Version 8, updated 3 June 2019. Available from http://hemiptera-databases.org/flow/ [accessed 15 Jan. 2020].
Bourgoin T., Wang R.R., Asche M., Hoch H., Soulier-Perkins A., Stroiński A., Yap S. & Szwedo J. 2015. From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology 134 (1): 63–77. https://doi.org/10.1007/s00435-014-0243-6
Bourgoin T., Guilbert E. & Wang M. 2018. Issidae molecular K-libration. In: 8th European Hemiptera Congress and 11th International Workshop on Leafhoppers and Planthoppers of Economic Importance, 24–29 June 2018: 36. University of Silesia, Zawiercie, Poland.
Gnezdilov V.M. 2013a. “Issidisation” of fulgoroid planthoppers (Homoptera, Fulgoroidea) as a case of parallel adaptive radiation. Entomological Review 93: 825–830. https://doi.org/10.1134/S001387381307004X
Gnezdilov V.M. 2013b. A modern classification of the family Caliscelidae Amyot et Serville (Homoptera, Fulgoroidea). Zoologicheskii Zhurnal 92: 1309–1311. [English translation in Entomological Review 2014 94: 211–214.] https://doi.org/10.1134/S0013873814020092
Gnezdilov V.M. & Bourgoin T. 2016. On the taxonomic position of Issus reticulatus Bervoets, 1910 (Hemiptera: Fulgoroidea: Issidae) from Baltic amber. Entomological Review 96 (5): 631–633. https://doi.org/10.1134/S0013873816050092
Hellmund M. 2018. The former Geiseltal Museum (1934–2011), the Eocene Geiseltal Fossilagerstätte (Germany) and the scientific meaning of Ben Barnes as a pioneer of systematic quantitative vertebrate excavations in the Geiseltal Lignites. Anuário do Instituto de Geociências – UFRJ 41 (1): 108–119. https://doi.org/10.11137/2018_1_108_119
Huys R., Suárez-Morales E., Serrano-Sánchez M. de Lourdes, Centeno-García E. & Vega F.J. 2016. Early Miocene amber inclusions from Mexico reveal antiquity of mangrove-associated copepods. Scientific Reports 6: 34872. https://doi.org/10.1038/srep34872
Kedves M. & Russell D.E. 1982. Palynology of the Thanetian layers of Menat. The geology of the Menat Basin, France. Palaeontographica, Abteilung В 182: 87–150.
Mayr G., Hervet S. & Buffetaut E. 2019. On the diverse and widely ignored Paleocene avifauna of Menat (Puy-de-Dôme, France): new taxonomic records and unusual soft tissue preservation. Geological Magazine 156 (3): 572–584. https://doi.org/10.1017/S0016756818000080
McKellar R.C., Wolfe A.P., Tappert R. & Muehlenbachs K. 2008. Correlation of Grassy Lake and Cedar Lake ambers using infrared spectroscopy, stable isotopes, and palaeoentomology. Canadian Journal of Earth Sciences 45: 1061–1082. https://doi.org/10.1139/E08-049
PaleoBioDB 2019. Paleodatabase. Available from https://paleobiodb.org/classic/displayStrata?geo
logical_group=&formation=Menat&group_formation_member=Menat [accessed 21 Aug. 2019].
Sadowski E.M., Schmidt A.R., Seyfullah L.J. & Kunzmann L. 2017. Conifers of the ‘Baltic amber forest’ and their palaeoecological significance. Stapfia 106: 1–73.
Shcherbakov D.E. 1996. Origin and evolution of the Auchenorhyncha as shown by the fossil record. In: Schaefer C.W. (ed.) Studies on Hemipteran Phylogeny: 31–45. Thomas Say Publications in Entomology, Lanham, Maryland.
Skidmore R.E. 1999. Checklist of Canadian Amber Inclusions in the Canadian National Collection of Insects. Research Branch Agriculture and Agri-Food Canada, electronic publication, but no longer available.
Steinheimer F.D. & Hastings K.A. 2018. Halle: the Geiseltal collection of Martin Luther University, Halle-Wittenberg. In: Beck L.A. & Joger U. (eds) Paleontological Collections of Germany, Austria and Switzerland: the History of Life of Fossil Organisms at Museums and Universities: 271–280. Natural History Collections, Springer, Cham. https://doi.org/10.1007/978-3-319-77401-5_25
Stroiński A. & Szwedo J. 2008. Thionia douglundbergi sp. nov. from the Miocene Dominican amber (Hemiptera: Fulgoromorpha: Issidae) with notes on extinct higher planthoppers. Annales Zoologici 58 (3): 529–536. https://doi.org/10.3161/000345408X364355
Stroiński A. & Szwedo J. 2012. The oldest known Lophopidae planthopper (Hemiptera: Fulgoromorpha) from the European Palaeocene. Geobios 45: 413–420. https://doi.org/10.1016/j.geobios.2011.10.007
Szwedo J., Bourgoin T. & Lefebvre F. 2006. New Mnemosynini taxa (Hemiptera, Fulgoromorpha: Cixiidae) from the Palaeogene of France with notes on their early association with host plants. Zootaxa 1122: 25–45.
Szwedo J., Drohojowska J., Popov Y.A., Simon E. & Węgierek P. 2019. Aphids, true hoppers, jumping plant-lice, scale insects, true bugs and whiteflies (Insecta: Hemiptera) from the Insect Limestone (latest Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh. https://doi.org/10.1017/S175569101900001X [published online 9 Aug. 2019].
Wang M., Zhang Y.L. & Bourgoin T. 2016. Planthopper family Issidae (Insecta: Hemiptera: Fulgoromorpha): linking molecular phylogeny with classification. Molecular Phylogenetics and Evolution 105: 224–234. https://doi.org/10.1016/j.ympev.2016.08.012
Wappler T., Currano E.D., Wilf P., Rust J. & Labandeira C.C. 2009. No post-Cretaceous ecosystem depression in European forests? Rich insect feeding damage on diverse middle Palaeocene plants, Menat, France. Proceedings of the Royal Society of London B: Biological Sciences 276: 4271–4277. https://doi.org/10.1098/rspb.2009.1255
Wedmann S., Uhl D., Lehmann T., Garrouste R., Nel A., Gomez B., Smith K. & Schaal S.F.K. 2018. The Konservat-Lagerstätte Menat (Paleocene; France) – an overview and new insights. Geologica Acta 16 (2): 189–213. https://doi.org/10.1344/GeologicaActa2018.16.2.5
Weitschat W. & Wichard W. 2010. Baltic Amber. In: Penney D. (ed.) Biodiversity of Fossils in Amber from the Major World Deposits: 80–115. Siri Scientific Press, Manchester.
Zhao S., Bourgoin T. & Wang M. 2019. The impact of a new genus on the molecular phylogeny of Hemisphaeriini (Hemiptera, Fulgoromorpha, Issidae). ZooKeys 880: 61–74.
Copyright (c) 2020 Thierry Bourgoin, Menglin Wang, André Nel

This work is licensed under a Creative Commons Attribution 4.0 International License.
Creative Commons Copyright Notices
Authors who publish with this journal agree to the following terms:
- Authors retain copyright and grant the journal right of first publication with the work simultaneously licensed under a Creative Commons Attribution License (CC BY 4.0) that allows others to share the work with an acknowledgement of the work's authorship and initial publication in this journal.
- Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.
- Authors are NOT PERMITTED to post their submitted work online (e.g., in institutional repositories or on personal websites) prior to publication, as it may lead to nomenclatural issues.